 In the third episode of the series, the narrator states that Yossarian was once Flower's mate and the dominant male of the Whiskers, but Zaphod dethroned him before the series started.The "" film states that Yossarian and Zaphod were once roving Lazuli males.Yossarian became the dominant male of the Whiskers when Flower's sister was the dominant female, and then left the group when Flower assumed the role.