 and concluded that the "Hysteriaceae" possess the pseudoparaphysate Pleospora-type centrum (all the structures enclosed within the ascocarp), in which cellular, septate pseudoparaphyses grow downwards from the cavity roof, initially anchored at both ends, and occupy the locule prior to the formation of asci.Even though "G. stellatum" clearly possessed the Pleospora-type centrum, it differed in that the locule was an elongated hystereothecium, and the locule appeared to be formed largely by the dissolution of the sterile centrum parenchyma, as in the Dothidea-developmental type.Luttrell (1953) thus concluded that locule formation in "G. stellatum", presumably representative of the Hysteriaceae as a whole, is intermediate between the "Pleospora" and "Dothidea" types.