 The Hysteriales were placed in the subclass Loculoascomycetes by Luttrell (1955), corresponding to the Ascoloculares first proposed by Nannfeldt (1932).Unlike the hymenoascomycetes, the loculoascomycete ascoma originates prior to karyogamy in the dikaryon, with the correlated character state being the functionally two-walled ascus which ruptures in a fissitunicate (like a Jack-in-the-box) fashion.Luttrell (1951) studied ascomal ontogeny and hamathicial development in "Glonium stellatum" Mühlenb.