 It begins with the first appearance of the ammonites "Drepanites rutherfordi" and "Cyrtopleuritis bicrenatus" and with the conodont "Cypridodella multidentata".The stage ends with the first appearance of the ammonites "Gnomohalorites cordilleranus" and "Sagenites quinquepunctatus" and the conodont "Cypridodella bidentata".The holotype MPUM 6009 of the "Bergamodactylus" pterosaur was found in a layer of the Zorzino Limestone Formation dating from the early Norian (upper Alaunian).