 His brother William would go on to become chief forester of the U.S. Forest Service.He graduated from Stanford University in 1898, and spent one year as a graduate student in zoology, during which he went to Alaska with the fur-seal expedition and to Brazil with the Banner-Agassiz expedition, where he made most of the biological collections.The following year he was an instructor at San Diego Normal School (now San Diego State University), leaving there to enter the University of Chicago as fellow in physiology.