 There he made a series of significant contributions to our understanding of the exocytic and endocytic pathways in neuroendocrine and exocrine cells and the exploitation of these pathways by enveloped animal viruses.He also provided the first evidence that endocytic membranes are used in the morphogenesis of vaccinia and human cytomegalovirus.Beginning in 1966 Tooze had been writing under the byline “our cell biology correspondent” a regular weekly column for the news and views section of the scientific journal Nature.