 Other major figures in the synthesis included E. B. Ford, Bernhard Rensch, Ivan Schmalhausen, and George Gaylord Simpson.An early event in the modern synthesis was R. A. Fisher's 1918 paper on mathematical population genetics, but William Bateson, and separately Udny Yule, were already starting to show how Mendelian genetics could work in evolution in 1902.Different syntheses followed, accompanying the gradual breakup of the early 20th century synthesis, including with social behaviour in E. O. Wilson's sociobiology in 1975, evolutionary developmental biology's integration of embryology with genetics and evolution, starting in 1977, and Massimo Pigliucci's proposed extended evolutionary synthesis of 2007.