 The biometricians argued vigorously against mutationism, saying that empirical evidence indicated that variation was continuous in most organisms, not discrete as Mendelism seemed to predict; they wrongly believed that Mendelism inevitably implied evolution in discontinuous jumps.A traditional view is that the biometricians and the Mendelians rejected natural selection and argued for their separate theories for 20 years, the debate only resolved by the development of population genetics.A more recent view is that Bateson, de Vries, Thomas Hunt Morgan and Reginald Punnett had by 1918 formed a synthesis of Mendelism and mutationism.