 In 1307, both were reunited by Nicholas II, but they were divided again in 1316 into Werle-Güstrow and Werle-Goldberg.In 1337, Werle-Waren split off from the Werle-Güstrow line.With the death of the William, the last lord on September 7, 1436, the fiefdom once again reverted to Mecklenburg.