 Villabruna 1 belonged to the Epigravettian culture.R-L389, also known as R1b1a1 (L388/PF6468, L389/PF6531), contains the very common subclade R-P297 and the rare subclade R-V1636.It is unknown whether all previously reported R-L389*(xP297) belong to R-V1636 or not.