 The ecological footprint of Croatia's population and industry varies significantly between the country's regions since 50% of the population resides in 26.8% of the nation's territory, with a particularly high impact made by the city of Zagreb and Zagreb County areas—their combined area comprises 6.6% of Croatia's territory while encompassing 25% of the population.The ecological footprint is most notably from the increased development of settlements and the sea coast leading to habitat fragmentation.Between 1998 and 2008, the greatest changes of land use pertained to artificially developed areas, but the scale of development is negligible compared to EU member states.