 The rise of eukaryotic cells at around 2.7 Gya, the rise of multicellular organisms at about 2 Gya, and the rise of mobile predators (around 600 Mya – 2 Gya, probably around 1 Gya) have all been attributed to early predatory behavior, and many very early remains show evidence of boreholes or other markings attributed to small predator species.Among more familiar species, the first vertebrate carnivores were fish, and then amphibians that moved on to land.Early tetrapods were large amphibious piscivores.