 He gained a reputation as an ingenious experimenter, as well as working to revise established medical textbooks.From 1927 to 1947 he was professor of physiology at St Bartholomew's Hospital, and from 1947 to 1951 director of the vision research unit of the Medical Research Council.He was president of the Quekett Microscopical Club from 1951 to 1954 and he was elected an Honorary Member in 1952.