 Route 207 originates from London United Tramways route 7 which ran between Uxbridge and Shepherd's Bush Green.In 1936, this route was replaced by a trolleybus route 607 which ran until 1960.Route 207 was introduced on 9 November 1960 as part of the trolleybus replacement program stage 8, replacing trolleybus route 607 between Uxbridge and Shepherd's Bush Green, using AEC Routemasters.