 The dominant females are the principal egg layers, while the subordinate females ("auxiliaries") or workers primarily forage and do not lay eggs.This hierarchy is not permanent, though; when the queen is removed from the nest, the second-most dominant female takes over the role of the previous queen.Dominance in females is determined by the severity of the scatteredness in the coloration of the clypeus (face), whereas dominance in males is shown by the variation of spots of their abdomens.