totally plastic. In 2002, a paper by Sparks and Jantz re-evaluated some of Boas's original data using new statistical techniques and concluded that there was a "relatively high genetic component" of head shape. Ralph Holloway of Columbia University argues that the new research raises questions about whether the variations in skull shape have "adaptive meaning and whether, in fact, normalizing selection might be at work on the trait, where both extremes, hyperdolichocephaly and hyperbrachycephaly, are at a slight selective disadvantage." In 2003, anthropologists Clarence C. Gravlee, H. Russell Bernard, and William R. Leonard reanalyzed Boas's data and concluded that most of Boas's original findings were correct. Moreover, they applied new statistical, computer-assisted methods to Boas's data and discovered more evidence for cranial plasticity. In a later publication, Gravlee, Bernard and Leonard reviewed Sparks's and Jantz's analysis. They argue that Sparks and Jantz misrepresented Boas's claims, and that Sparks's and Jantz's data support Boas. For example, they point out that Sparks and Jantz look at changes in cranial size in relation to how long an individual has been in the United States in order to test the influence of the environment. Boas, however, looked at changes in cranial size in relation to how long the mother had been in the United States. They argue that Boas's method is more useful, because the prenatal environment is a crucial developmental factor. Jantz and Sparks responded to Gravlee et al., reiterating that Boas' findings lacked biological meaning, and that the interpretation of Boas' results common in the literature was biologically inaccurate. In a later study, the same authors concluded that the effects Boas observed were likely the result of population-specific environmental effects such as changes in cultural practices for cradling infants, rather than the effects of a general "American environment" which caused populations in America to converge to a common cranial type, as Boas had suggested. Modern use in animal breeding The cephalic index is used in the categorisation of animals, especially breeds of dogs and cats. Brachycephalic animals A brachycephalic skull is relatively broad and short (typically with the breadth at least 80% of the length). Dog breeds such as the pug are sometimes classified as "extreme brachycephalic". Because of the health issues brachycephaly is regarded as torture breeding as it often leads to the brachycephalic airway obstructive syndrome. List of brachycephalic dogs Affenpinscher American Bulldog American Bully Boston Terrier Boxer Brussels Griffon