genealogy and similarity count in classification (in a manner determined by the evolutionary taxonomist). It is among the cladists that Occam's razor is applied, through the method of cladistic parsimony. Cladistic parsimony (or maximum parsimony) is a method of phylogenetic inference that yields phylogenetic trees (more specifically, cladograms). Cladograms are branching, diagrams used to represent hypotheses of relative degree of relationship, based on synapomorphies. Cladistic parsimony is used to select as the preferred hypothesis of relationships the cladogram that requires the fewest implied character state transformations (or smallest weight, if characters are differentially weighted). Critics of the cladistic approach often observe that for some types of data, parsimony could produce the wrong results, regardless of how much data is collected (this is called statistical inconsistency, or long branch attraction). However, this criticism is also potentially true for any type of phylogenetic inference, unless the model used to estimate the tree reflects the way that evolution actually happened. Because this information is not empirically accessible, the criticism of statistical inconsistency against parsimony holds no force. For a book-length treatment of cladistic parsimony, see Elliott Sober's Reconstructing the Past: Parsimony, Evolution, and Inference (1988). For a discussion of both uses of Occam's razor in biology, see Sober's article "Let's Razor Ockham's Razor" (1990). Other methods for inferring evolutionary relationships use parsimony in a more general way. Likelihood methods for phylogeny use parsimony as they do for all likelihood tests, with hypotheses requiring fewer differing parameters (i.e., numbers or different rates of character change or different frequencies of character state transitions) being treated as null hypotheses relative to hypotheses requiring more differing parameters. Thus, complex hypotheses must predict data much better than do simple hypotheses before researchers reject the simple hypotheses. Recent advances employ information theory, a close cousin of likelihood, which uses Occam's razor in the same way. Of course, the choice of the "shortest tree" relative to a not-so-short tree under any optimality criterion (smallest distance, fewest steps, or maximum likelihood) is always based on parsimony Francis Crick has commented on potential limitations of Occam's razor in biology. He advances the argument that because biological systems are the products of (an ongoing) natural selection, the mechanisms are not necessarily optimal in an obvious sense. He cautions: "While Ockham's razor is a useful tool in the physical sciences, it can be a very dangerous implement in biology. It is thus very rash