Archaic periods, such as the Tomb of the Augurs, the Tomb of the Triclinium or the Tomb of the Leopards, as well as other tombs from the archaic period in the Monterozzi necropolis in Tarquinia, were painted by Greek painters or, in any case, foreigner artists. These images have, therefore, a very limited value for a realistic representation of the Etruscan population. It was only from the end of the 4th century B.C. that evidence of physiognomic portraits began to be found in Etruscan art and Etruscan portraiture became more realistic. Genetic research An mtDNA study in 2004, based on Etruscan samples from Veneto, Tuscany, Lazio, and Campania, stated that the Etruscans had no significant heterogeneity, and that all mitochondrial lineages observed among the Etruscan samples appear typically European, but only a few haplotypes were shared with modern populations. Allele sharing between the Etruscans and modern populations is highest among Germans (seven haplotypes in common), the Cornish from South West England (five haplotypes in common), the Turks (four haplotypes in common), and the Tuscans (two haplotypes in common). A couple of mitochondrial DNA studies, published in 2013 in the journals PLOS One and American Journal of Physical Anthropology, based on Etruscan samples from Tuscany and Latium, concluded that the Etruscans were an indigenous population, showing that Etruscan mtDNA appears to fall very close to a Neolithic population from Central Europe (Germany, Austria, Hungary) and to other Tuscan populations, strongly suggesting that the Etruscan civilization developed locally from the Villanovan culture, as already supported by archaeological evidence and anthropological research, and that genetic links between Tuscany and western Anatolia date back to at least 5,000 years ago during the Neolithic and the "most likely separation time between Tuscany and Western Anatolia falls around 7,600 years ago", at the time of the migrations of Early European Farmers (EEF) from Anatolia to Europe in the early Neolithic. The ancient Etruscan samples had mitochondrial DNA haplogroups (mtDNA) JT (subclades of J and T) and U5, with a minority of mtDNA H1b. In the collective volume Etruscology published in 2017, British archeologist Phil Perkins provides an analysis of the state of DNA studies and writes that "none of the DNA studies to date conclusively prove that Etruscans were an intrusive population in Italy that originated in the Eastern Mediterranean or Anatolia" and "there are indications that the evidence of DNA can support the theory that