Naturae. The genus name Panthera was coined by Lorenz Oken in 1816. Between the mid-18th and mid-20th centuries, 26 lion specimens were described and proposed as subspecies, of which 11 were recognised as valid in 2005. They were distinguished mostly by the size and colour of their manes and skins. Subspecies In the 19th and 20th centuries, several lion type specimens were described and proposed as subspecies, with about a dozen recognised as valid taxa until 2017. Between 2008 and 2016, IUCN Red List assessors used only two subspecific names: P. l. leo for African lion populations, and P. l. persica for the Asiatic lion population. In 2017, the Cat Classification Task Force of the Cat Specialist Group revised lion taxonomy, and recognises two subspecies based on results of several phylogeographic studies on lion evolution, namely: P. l. leo − the nominate lion subspecies includes the Asiatic lion, the regionally extinct Barbary lion, and lion populations in West and northern parts of Central Africa. Synonyms include P. l. persica , P. l. senegalensis , P. l. kamptzi , and P. l. azandica . Multiple authors referred to it as 'northern lion' and 'northern subspecies'. P. l. melanochaita − includes the extinct Cape lion and lion populations in East and Southern African regions. Synonyms include P. l. somaliensis , P. l. massaica , P. l. sabakiensis , P. l. bleyenberghi , P. l. roosevelti , P. l. nyanzae , P. l. hollisteri , P. l. krugeri , P. l. vernayi , and P. l. webbiensis . It has been referred to as 'southern subspecies' and 'southern lion'. However, there seems to be some degree of overlap between both groups in northern Central Africa. DNA analysis from a more recent study indicates, that Central African lions are derived from both northern and southern lions, as they cluster with P. leo leo in mtDNA-based phylogenies whereas their genomic DNA indicates a closer relationship with P. leo melanochaita. Lion samples from some parts of the Ethiopian Highlands cluster genetically with those from Cameroon and Chad, while lions from other areas of Ethiopia cluster with samples from East Africa. Researchers therefore assume Ethiopia is a contact zone between the two subspecies. Genome-wide data of a wild-born historical lion sample from Sudan showed that it clustered with P. l. leo in mtDNA-based phylogenies, but with a high affinity to P. l. melanochaita. This result suggested that