ancestry could be traced to the Middle East, from where it was introduced to Europe by returning Crusaders. Little external evidence supports this theory. Another theory stated that it originated from an ancient indigenous vine found in Cyprus. Modern DNA fingerprinting research at University of California, Davis, now suggests that Chardonnay is the result of a cross between the Pinot noir and Gouais blanc (Heunisch) grape varieties. The Romans are thought to have brought Gouais blanc from Croatia, and it was widely cultivated by peasants in eastern France. The Pinot of the French aristocracy grew in close proximity to the Gouais blanc, giving the two ample opportunity to interbreed. Since the two parents were genetically distant, many of the crosses showed hybrid vigour and were selected for further propagation. These "successful" crosses included Chardonnay and siblings such as Aligoté, Aubin vert, Auxerrois, Bachet noir, Beaunoir, Franc Noir de la-Haute-Saône, Gamay Blanc Gloriod, Gamay noir, Melon, Knipperlé, Peurion, Roublot, Sacy, and Dameron. Clones, crossing, and mutations As of 2006, 34 clonal varieties of Chardonnay could be found in vineyards throughout France; most of these were developed at the University of Burgundy in Dijon. The so-called "Dijon clones" are bred for their adaptive attributes, with vineyard owners planting the clonal variety best suited to their terroir and which will produce the characteristics that they are seeking in the wine. Examples include the lower-yielding clones 'Dijon-76', '95' and '96' that produce more flavor-concentrated clusters. 'Dijon-77' and '809' produce more aromatic wines with a "grapey" perfume, while 'Dijon-75', '78', '121', '124', '125' and '277' are more vigorous and higher-yielding clones. New World varieties include the 'Mendoza' clone, which produced some of the early California Chardonnays. The 'Mendoza' clone is prone to develop millerandage, also known as "hens and chicks", where the berries develop unevenly. In places such as Oregon, the use of newer Dijon clones has had some success in those regions of the Willamette Valley with climates similar to that of Burgundy. Chardonnay has served as parent to several French-American hybrid grapes, as well as crossings with other V. vinifera varieties. Examples include the hybrid Chardonel, which was a Chardonnay and Seyval blanc cross produced in 1953 at the New York State Agricultural Experiment Station. Mutations of the Chardonnay grape include the rare pink-berried 'Chardonnay Rose'; also 'Chardonnay Blanc Musqué', which produces an intensely aromatic wine. Chardonnay Blanc Musqué is mostly found around