egg to its first birthday is only 19%. For females, the annual survival rate rises to 45% for juveniles and 95% for adults. The male survival rates follow a similar pattern, but are probably lower overall than females, as evidenced by the average male age being lower than that of the female. Natural disasters can confound age distributions. For instance, a hurricane can destroy many nests in a region, resulting in fewer hatchlings the next year. Age distributions may also be skewed by migrations of adults. To understand painted turtle adult age distributions, researchers require reliable methods. Turtles younger than four years (up to 12 years in some populations) can be aged based on "growth rings" in their shells. For older turtles, some attempts have been made to determine age based on size and shape of their shells or legs using mathematical models, but this method is more uncertain. The most reliable method to study the long-lived turtles is to capture them, permanently mark their shells by notching with a drill, release the turtles, and then recapture them in later years. The longest-running study, in Michigan, has shown that painted turtles can live more than 55 years. Adult sex ratios of painted turtle populations average around 1:1. Many populations are slightly male-heavy, but some are strongly female-imbalanced; one population in Ontario has a female to male ratio of 4:1. Hatchling sex ratio varies based on egg temperature. During the middle third of incubation, temperatures of produce males, and anything above or below that, females. It does not appear that females choose nesting sites to influence the sex of the hatchlings; within a population, nests will vary sufficiently to give both male and female-heavy broods. Ecology Diet The painted turtle is a bottom-dwelling hunter. It quickly juts its head into and out of vegetation to stir potential victims out into the open water, where they are pursued. Large prey is ripped apart with the forefeet as the turtle holds it in its mouth. It also consumes plants and skims the surface of the water with its mouth open to catch small particles of food. Although all subspecies of painted turtle eat both plants and animals (in the form of leaves, algae, fish, crustaceans, aquatic insects and carrion), their specific diets vary. Painted turtles obtain coloration from carotenoids in their natural diet by eating algae and a variety of aquatic plants from