tools. They were only retouching larger flakes, the fourteen such tools averaging : one marginally retouched flake, one notch, three spines, seven denticulate sidescrapers, and one denticulate point. Fire and palaeoclimate Only a few charcoal particles have been collected from TD6, which probably originated from a fire well outside the cave. There is no evidence of any fire use or burnt bones (cooking) in the occupation sequences of the Gran Dolina. In other parts of the world, reliable evidence of fire usage does not surface in the archaeological record until roughly 400,000 years ago. In 2016, small mammal bones burned in fires exceeding were identified from 780- to 980-thousand-year-old deposits at in southern Spain, which potentially could have come from a human source as such a high temperature is usually (though not always) recorded in campfires as opposed to natural bushfires. Instead of using fire, these early Europeans probably physiologically withstood the cold, such as by eating a high protein diet to support a heightened metabolism. Despite glacial cycles, the climate was probably similar or a few degrees warmer compared to that of today's, with the coldest average temperature reaching sometime in December and January, and the hottest in July and August . Freezing temperatures could have been reached from November to March, but the presence of olive and oak suggests subfreezing was an infrequent occurrence. TE9 similarly indicates a generally warm climate. The Happisburgh footprints were lain in estuarine mudflats with open forests dominated by pine, spruce, birch, and in wetter areas alder, with patches of heath and grasslands; the vegetation is consistent with the cooler beginning or end of an interglacial. H. antecessor probably migrated from the Mediterranean shore into inland Iberia when colder glacial periods were transitioning to warmer interglacials, and warm grasslands dominated, vacating the region at any other time. They may have followed water bodies while migrating, in the case of Sierra de Atapuerca, most likely the Ebro River. Food The fossils of sixteen animal species were recovered randomly mixed with the H. antecessor material at the Gran Dolina, including the extinct bush-antlered deer, the extinct species of fallow deer Dama vallonetensi, the extinct subspecies of red deer Cervus elaphus acoronatus, the extinct bison Bison voigstedtensi, the extinct rhino Stephanorhinus etruscus, the extinct horse Equus stenonis, the extinct fox Vulpes praeglacialis, the extinct bear Ursus dolinensis, the extinct wolf Canis mosbachensis, the spotted hyena, the