– and the surrounding region was also well inhabited, to judge from over 50 villas discovered in the area. It was mentioned by the 1st century AD geographer Pomponius Mela, who described it in his work De situ orbis libri III as one of "the wealthiest cities, albeit the wealthiest among small ones" in Mauretania. It is also mentioned by Pliny the Elder, and the 2nd century Antonine Itinerary refers to its location and names it as Volubilis Colonia. Its population was dominated by Romanised Berbers. The city remained loyal to Rome despite a revolt in 40–44 AD led by one of Ptolemy's freedmen, Aedemon, and its inhabitants were rewarded with grants of citizenship and a ten-year exemption from taxes. The city was raised to the status of a municipium and its system of governance was overhauled, with the Punic-style suffetes replaced by annually elected duumvirs, or pairs of magistrates. However, the city's position was always tenuous; it was located on the south-eastern edge of the province, facing hostile and increasingly powerful Berber tribes. A ring of five forts located at the modern hamlets of Aïn Schkor, Bled el Gaada, Sidi Moussa, Sidi Said and Bled Takourart (ancient Tocolosida) were constructed to bolster the city's defence. Sidi Said was the base for the Cohors IV Gallorum equitata, an auxiliary cavalry unit from Gaul, while Aïn Schkor housed Hispanic and Belgic cohorts. Sidi Moussa was the location of a cohort of Parthians, and Gallic and Syrian cavalry were based at Toscolosida. Rising tensions in the region near the end of the 2nd century led the emperor Marcus Aurelius to order the construction of a circuit of walls with eight gates and 40 towers. Volubilis was connected by road to Lixus and Tingis (capital city of the Roman province of Mauretania Tingitana, modern Tangier) but had no eastwards connections with the neighbouring province of Mauretania Caesariensis, as the territory of the Berber Baquates tribe lay in between. A Jewish community existed in Volubilis in the third century, as evident from several Hebrew, Greek and Latin funeral inscriptions and Menorah-shaped lamps. It is the most south-western location where ancient Hebrew inscription has been found. Rome's control over the city ended following the chaos of the Crisis of the Third Century, when the empire nearly disintegrated as a series of generals seized and lost power through civil wars, palace coups and assassinations. Around 280,