The biodiversity of New Caledonia is of exceptional biological and paleoecological interest. It is frequently referred to as a biodiversity hotspot. The country is a large South Pacific archipelago with a total land area of more than . The terrain includes a variety of reefs, atolls, small islands, and a variety of topographical and edaphic regions on the largest island, all of which promote the development of unusually concentrated biodiversity. The region's climate is oceanic and tropical. New Caledonia is separated from the nearest mainland by more than of open sea. Its isolation dates from at least the mid-Miocene, and possibly from the Oligocene, and that isolation has preserved its relict biota, fostering the evolution of wide ranges of endemic species. Location and description New Caledonia lies on the southernmost edge of the tropical zone, near the Tropic of Capricorn. It is part of the Melanesia subregion. It includes the main island of Grande Terre, the Loyalty Islands, the Belep archipelago, the Isle of Pines, the Chesterfield Islands in the Coral Sea, and a few remote islets. The archipelago is about east of Australia and 1500 km, 1800 km and 1200 km from New Zealand, Papua New Guinea and Fiji respectively. A few lesser islands are closer, but now provide no convenient island-hopping path by which animal species could pass either to or from major mainlands. Some plants, invertebrates, sea mammals, and many flying species as aquatic birds, parrots, and bats have spread to new locations, either under their own power or due to freak events such as storms, or have been transported by humans. Some plant species have colonized new areas by means of seed carried by ocean currents. Some animal and plant species have reached New Caledonia from surrounding regions and in turn, some New Caledonian species have successfully extended their ranges into the Pacific Ocean area. Other New Caledonian species or their close relatives are found in regions remote from the archipelago. For example, the New Caledonian parakeet is known to be the basal species in the genus Cyanoramphus, which has spread to many Pacific islands. Many bats and birds that rely heavily on fruit for their diet, including members of the families Cotingidae, Columbidae, Trogonidae, Turdidae, and Toucan, swallow seeds, then regurgitate them or pass them in their faeces. Such seed dispersal (ornithochory) has been a major mechanism of seed dispersal across ocean barriers. Seeds of