under constant surveillance by their dam, while the sire and elder offspring provide food. The pups open their eyes after 8–10 days and emerge from the den at the age of 3 weeks. They are weaned at 8–9 weeks, and can hunt by themselves at the age of 6 months. Sexual maturity is attained at 11 months, though few black-backed jackals reproduce in their first year. Unlike golden jackals, which have comparatively amicable intrapack relationships, black-backed jackal pups become increasingly quarrelsome as they age, and establish more rigid dominance hierarchies. Dominant pups appropriate food, and become independent at an earlier age. The grown pups may disperse at one year of age, though some remain in their natal territories to assist their parents in raising the next generation of pups. The average lifespan in the wild is 7 years, though captive specimens can live twice as long. Ecology Habitat The species generally shows a preference for open areas with little dense vegetation, though it occupies a wide range of habitats, from arid coastal deserts to areas with more than 2000 mm of rainfall. It also occurs in farmlands, savannas, open savanna mosaics, and alpine areas. Diet Black-backed jackals are omnivores, which feed on invertebrates, such as beetles, grasshoppers, crickets, termites, millipedes, spiders, and scorpions. They also feed on mammals, such as rodents, hares, and young antelopes up to the size of topi calves. They also feed on carrion, birds, lizards and snakes. A pair of black-backed jackals in the Kalahari desert was observed to kill a kori bustard, and on a separate occasion, a black mamba by prolonged harassment of the snake and crushing of the snake's head. Black-backed jackals occasionally feed on fruits and berries. It also feeds on eggs of birds. In coastal areas, they feed on beached marine mammals, seals, fish, and mussels. A single jackal is capable of killing a healthy adult impala. Adult dik-diks and Thomson's gazelles seem to be the upper limit of their killing capacity, though they target larger species if those are sick, with one pair having been observed to harass a crippled bull rhinoceros. They typically kill tall prey by biting at the legs and loins, and frequently go for the throat. In Serengeti woodlands, they feed heavily on African grass rats. In East Africa, during the dry season, they hunt the young of gazelles, impalas, topi, tsessebe, and warthogs. In South