in New Caledonia. Pancheria and Codia are endemic, although the last is known as a fossil in Australia, while Cunonia has 23 endemic species in New Caledonia and one species in South Africa. The other three genera have a Papuan-Australian (Acsmithia), Australian (Geissois) and sub-Antarctic (Weinmannia) distribution. Proteaceae's two main centers of dispersion are in Australia and Southern Africa; the family is represented in the archipelago by 43 species. They divide into six endemic genera: Beauprea, Beaupreopsis, Garnieria, Kermadecia, Sleumerodendron and Virotia and three non-endemic genera Grevillea, Knightia and Stenocarpus. Myrtaceae, although basically Gondwanan in origin, currently range throughout the tropical world and have reached the Northern Hemisphere. With 229 species, it is the largest family in New Caledonia. It has two widely distributed genera, Eugenia and Syzygium. The endemic genera are Arillastrum, Cloezia, Myrtastrum, Pleurocalyptus, and Purpureostemon. The genus Melaleuca is represented by six endemic species and one shared with Australia and Papua New Guinea, Melaleuca quinquenervia, known locally as niaouli. Fauna The New Caledonia Great Barrier Reef is a UNESCO World Heritage Site and the second largest barrier reef in the world. Amedee island is a special marine reserve of coral reef lagoon, Ilot aux Goelands is a tiny lagoon island surrounded by a large shallow reef flat. The reef has great species diversity with a high level of endemism. Many groups have been under sampled and insufficiently studied, especially when considering hard bottoms of the intermediate coral reefs and external slopes of the barrier reef. This diversity includes oceanic and continental reefs forming islands, atolls, uplifted reefs, immerged reefs, fringing reefs, barrier reefs, patch reefs and shallow or deep lagoons, is home to endangered dugongs (Dugong dugon) and is an important nesting site for green sea turtle (Chelonia mydas). New Caledonia has a remarkable marine fauna due to the abundance of relic organisms from the Mesozoic., for example some sponges of the Lithistideae and the Tetractinellideae which are considered living fossils due to their closeness to Cretaceous species. The endemic mollusc Nautilus macromphalus is one of only four Nautilus species known in the world, the only living group of cephalopods with an external shell.. This mollusc seems to be identical to its Paleozoic ancestor. Cephalodiscus graptolitoides, described in 1993, is also considered a living fossil member of the graptolites, previously thought to be extinct for over 300 million years. Today's New Caledonian marine biodiversity has been counted