also identified the referred material of C. iguidensis as belonging to Sigilmassasaurus (later referred to Spinosaurus sp.) and a non-carcharodontosaurine, and therefore chose to limit C. iguidensis to the holotype pending future research. Description Carcharodontosaurus was one of the longest and heaviest known carnivorous dinosaurs, with an enormous long skull and long, serrated teeth up to long. C. saharicus reached in length and approximately in body mass, while C. iguidensis reached in length and in body mass. Brain and inner ear In 2001, Hans C. E. Larsson published a description of the inner ear and endocranium of Carcharodontosaurus saharicus. Starting from the portion of the brain closest to the tip of the animal's snout is the forebrain, which is followed by the midbrain. The midbrain is angled downwards at a 45-degree angle and towards the rear of the animal. This is followed by the hindbrain, which is roughly parallel to the forebrain and forms a roughly 40-degree angle with the midbrain. Overall, the brain of C. saharicus would have been similar to that of a related dinosaur, Allosaurus fragilis. Larsson found that the ratio of the cerebrum to the volume of the brain overall in Carcharodontosaurus was typical for a non-avian reptile. Carcharodontosaurus also had a large optic nerve. The three semicircular canals of the inner ear of Carcharodontosaurus saharicus – when viewed from the side – had a subtriangular outline. This subtriangular inner ear configuration is present in Allosaurus, lizards, turtles, but not in birds. The semi-"circular" canals themselves were actually very linear, which explains the pointed silhouette. In life, the floccular lobe of the brain would have projected into the area surrounded by the semicircular canals, just like in other non-avian theropods, birds, and pterosaurs. Classification The following cladogram after Apesteguía et al., 2016, shows the placement of Carcharodontosaurus within Carcharodontosauridae. Paleobiology Feeding A study by Donald Henderson, the curator of dinosaurs at the Royal Tyrrell Museum suggests that Carcharodontosaurus was able to lift animals weighing a maximum of in its jaws based on the strength of its jaws, neck, and its center of mass. Pathology SGM-Din 1, a Carcharodontosaurus saharicus skull, has a circular puncture wound in the nasal and "an abnormal projection of bone on the antorbital rim." References External links Student Identifies Enormous New Dinosaur December 7 2007 from the Science daily Discovery Channel Videos: Monsters Resurrected: Biggest Killer Dino Carcharodontosaurids Late Cretaceous dinosaurs of