The is a zoo, managed by the Tokyo Metropolitan Government, and located in Taitō, Tokyo, Japan. It is Japan's oldest zoo, opened on March 20, 1882. It is served by Ueno Station, Keisei Ueno Station and Nezu Station, with convenient access from several public transportation networks (JR East, Tokyo Metro and Keisei Electric Railway). The Ueno Zoo Monorail, the first monorail in the country, connects the eastern and western parts of the grounds. The zoo is in Ueno Park, a large urban park that is home to museums, a small amusement park, and other attractions. The zoo is closed on Mondays (Tuesday if Monday is a holiday). History The zoo started life as a menagerie attached to the National Museum of Natural History. In 1881, responsibility for this menagerie was handed to naturalist and civil servant Tanaka Yoshio, who oversaw its transition into a public zoo. The ground was originally estate of the imperial family, but was to the municipal government in 1924 — along with Ueno Park — on the occasion of crown prince Hirohito's wedding. World War II In August 1943, the administrator of Tokyo, Shigeo Ōdachi, ordered that all "wild and dangerous animals" at the zoo be killed, claiming that bombs could hit the zoo and escaped animals would wreak havoc in the streets of Tokyo. Requests by the staff at the zoo for a reprieve, or to evacuate the animals elsewhere, were refused. The animals were executed primarily by poisoning, strangulation or by simply placing the animals on starvation diets. A memorial service was held for the animals in September 1943 (while two of the elephants were still starving) and a permanent memorial (built anew in 1975) can be found in the Ueno Zoo. Shortly after the March 1945 bombings of Tokyo, the Japanese placed U.S. Army Air Force navigator and bombardier Ray "Hap" Halloran on display naked in a Ueno Zoo tiger cage so civilians could walk in front of the cage and view the B-29 prisoner. Recent renovations The zoo provides animals an environment similar to the natural habitat. In recent years, the old-fashioned cages of the past have been replaced with modern habitats, such as the "Gorilla Woods," built after two well-publicized mishaps in 1999. Animals The zoo is home to more than 2,600 individuals representing over 500 species. Principal animals After the death of giant panda Ling Ling in 2008, Ueno Zoo