The Roman conquest of Judea led by Pompey took place in 63 BC. The Roman occupation encompassed the end of Jewish independence in Judea, the last years of the Hasmonean kingdom, the Herodian age and the rise of Christianity, the First Jewish–Roman War, and the fall of Jerusalem and the destruction of the Second Temple. The total population of Pharisees, the forerunners of modern Rabbinic Judaism, was around 6,000 ("exakischilioi"), according to Josephus. Local population displacements occurred with the expulsion of the Jews from Jerusalem – "In the earlier revolt in the previous century, 66–73 CE, Rome destroyed the Temple and forbade Jews to live in the remaining parts of Jerusalem; for this reason, the Rabbis gathered instead on the Mediterranean coast in Yavneh near Jaffa". Dispersal to other parts of the Roman Empire occurred: "No date or origin can be assigned to the numerous settlements eventually known in the west, and some may have been founded as a result of the dispersal of Palestinian Jews after the revolts of AD 66–70 and 132-5, but it is reasonable to conjecture that many, such as the settlement in Puteoli attested in 4 BC, went back to the late republic or early empire and originated in voluntary emigration and the lure of trade and commerce." Modern estimates vary: Applebaum argues that in the Herodian kingdom, there were 1.5 million Jews, a figure Ben David says covers the numbers in Judea alone. Salo Wittmayer Baron estimated the population at 2.3 million at the time of Roman emperor Claudius (reigned 41–54). According to Israeli archeologist Magen Broshi, west of the Jordan River the population did not exceed 1 million: "... the population of Palestine in antiquity did not exceed a million persons. It can also be shown, moreover, that this was more or less the size of the population in the peak period – the late Byzantine period, around AD 600" Broshi made calculations based on the grain-producing capacity of Palestine and on its role in the indigenous diet, assuming an average annual per-capita consumption of 200 kg. (with a maximum of 250 kg.), which would work out to the limit of a sustainable population of 1,000,000 people, a figure which, Broshi states, remained roughly constant down to the end of the Byzantine period (600 CE). The proportion of Jews to gentiles is also unknown. Three events caused the Jewish population dominance to change