a run, would limit the speed of large theropods. Instead they posed that the imbalance caused by increasing velocity would be the limiting factor. Calculating the time it would take for a leg to gain balance after the retraction of the opposite leg, they found the upper kinematic limit of the running speed to be . They also found comparison between the running capability of Giganotosaurus and birds like the ostrich based on the strength of their leg-bones to be of limited value, since theropods, unlike birds, had heavy tails to counterbalance their weight. A 2017 biomechanical study of the running ability of Tyrannosaurus by the biologist William I. Sellers and colleagues suggested that skeletal loads were too great to have allowed adult individuals to run. The relatively long limbs, which were long argued to indicate good running ability, would instead have mechanically limited it to walking gaits, and it would therefore not have been a high-speed pursuit predator. They suggested that these findings would also apply to other long-limbed giant theropods such as Giganotosaurus, Mapusaurus, and Acrocanthosaurus. Feeding In 2002, Coria and Currie found that various features of the rear part of the skull (such as the frontwards slope of the occiput and low and wide occipital condyle) indicate that Giganotosaurus would have had a good capability of moving the skull sideways in relation to the front neck vertebrae. These features may also have been related to the increased mass and length of the jaw muscles; the jaw articulation of Giganotosaurus and other carcharodontosaurids was moved hindwards to increase the length of the jaw musculature, enabling faster closure of the jaws, whereas tyrannosaurs increased the mass of the lower jaw musculature, to increase the power of their bite. In 2005 Therrien and colleagues estimated the relative bite force of theropods and found that Giganotosaurus and related taxa had adaptations for capturing and bringing down prey by delivering powerful bites, whereas tyrannosaurs had adaptations for resisting torsional stress and crushing bones. Estimates in absolute values like newtons were impossible. The bite force of Giganotosaurus was weaker than that of Tyrannosaurus, and the force decreased hindwards along the tooth row. The lower jaws were adapted for slicing bites, and it probably captured and manipulated prey with the front part of the jaws. These authors suggested that Giganotosaurus and other allosaurs may have been generalized predators that fed on a wide spectrum