represented an animal of , if geometrically similar to the holotype specimen. By using multivariate regression equations, these authors also suggested an alternative weight of for the holotype and for the larger specimen, and that the latter was therefore the largest known terrestrial carnivore. In 2005, the paleontologist Cristiano Dal Sasso and colleagues described new skull material (a snout) of Spinosaurus (the original fossils of which were also destroyed during World War II), and concluded this dinosaur would have been long with a weight , exceeding the maximum size of all other theropods. In 2006, Coria and Currie described the large theropod Mapusaurus from Patagonia; it was closely related to Giganotosaurus and of approximately the same size. In 2007, the paleontologists François Therrien and Donald M. Henderson found that Giganotosaurus would have approached in length and in weight, while Carcharodontosaurus would have approached in length and in weight (surpassing Tyrannosaurus), and estimated the Giganotosaurus holotype skull to have been long. They cautioned that these measurements depended on whether the incomplete skulls of these animals had been reconstructed correctly, and that more complete specimens were needed for more accurate estimates. They also found that Dal Sasso and colleagues' reconstruction of Spinosaurus was too large, and instead estimated it to have been long, weighing , and possibly as low as in length and in weight. They concluded that these dinosaurs had reached the upper biomechanical size limit attainable by a strictly bipedal animal. In 2010, the paleontologist Gregory S. Paul suggested that the skulls of carcharodontosaurs had been reconstructed as too long in general. In 2012, the paleontologist Matthew T. Carrano and colleagues noted that though Giganotosaurus had received much attention due to its enormous size, and in spite of the holotype being relatively complete, it had not yet been described in detail, apart from the braincase. They pointed out that many contacts between skull bones were not preserved, which lead to the total length of the skull being ambiguous. They found instead that the skulls of Giganotosaurus and Carcharodontosaurus were exactly the same size as that of Tyrannosaurus. They also measured the femur of the Giganotosaurus holotype to be long, in contrast to the original measurement, and proposed that the body mass would have been smaller overall. In 2013, the paleontologist Scott Hartman published a Graphic Double Integration mass estimate (based on drawn skeletal reconstructions) on his blog, wherein he found