in epiphytes, especially orchids. Characteristic canopy trees are Cullenia exarillata, Mesua ferrea, Palaquium ellipticum, Gluta travancorica, and Nageia wallichiana. Nageia is a podocarp conifer with origins in the ancient supercontinent of Gondwana, of which India was formerly part, and a number of other plants in the ecoregion have Gondwana origins. Other evergreen tree species of the montane forest include Calophyllum austroindicum, Garcinia rubro-echinata, Garcinia travancorica, Diospyros barberi, Memecylon subramanii, Memecylon gracile, Goniothalamus rhynchantherus, and Vernonia travancorica. The other major habitat type in this ecoregion is the shola-grassland complex at elevations of . Shola is a stunted forest with small trees including Pygeum gardneri, Schefflera racemosa, Linociera ramiflora, Syzygium, Rhododendron nilgiricum, Mahonia napaulensis, Elaeocarpus recurvatus, Ilex denticulata, Michelia nilagirica, Actinodaphne bourdillonii and Litsea wightiana. The understorey consists of dense shrubs. These shola forest patches are interspersed with montane grasslands characterized by frost- and fire-resistant grass species like Chrysopogon zeylanicus, Cymbopogon flexuosus, Arundinella ciliata, Arundinella mesophylla, Arundinella tuberculata, Themeda tremula, and Sehima nervosum. Fauna Endemic fauna in this ecoregion include Nilgiri tahr (Nilgiritragus hylocrius), Nilgiri pipit (Anthus nilghiriensis), Nilgiri laughingthrush (Montecincla cachinnans), broad-tailed grassbird (Schoenicola platyurus), Nilgiri long-tailed tree mouse (Vandeleuria nilagirica), eight endemic reptile genera encompassing Brachyophidium, Dravidogecko, Melanophidium, Plectrurus, Ristella, Salea, Teretrurus and Xylophis with 90 species, and the six amphibian genera Indotyphlus, Melanobatrachus, Nannobatrachus, Nyctibatrachus, Ranixalus and Uraeotyphlus. The ecoregion also hosts Nilgiri langur (Semnopithecus johnii), Malabar large-spotted civet (Viverra civettina), brown palm civet (Paradoxurus jerdoni), Salim Ali's fruit bat (Latidens salimalii), Nilgiri striped squirrel (Funambulus sublineatus) and Layard's palm squirrel (F. layardi). References External links ecoregions of India Indomalayan ecoregions Montane forests Tropical and subtropical moist broadleaf forests Tropical rainforests of India