Macro photography (or photomacrography or macrography, and sometimes macrophotography) is extreme close-up photography, usually of very small subjects and living organisms like insects, in which the size of the subject in the photograph is greater than life size (though macrophotography also refers to the art of making very large photographs). By the original definition, a macro photograph is one in which the size of the subject on the negative or image sensor is life size or greater. In some senses, however, it refers to a finished photograph of a subject that is greater than life size. The ratio of the subject size on the film plane (or sensor plane) to the actual subject size is known as the reproduction ratio. Likewise, a macro lens is classically a lens capable of reproduction ratios of at least 1:1, although it often refers to any lens with a large reproduction ratio, despite rarely exceeding 1:1. Apart from technical photography and film-based processes, where the size of the image on the negative or image sensor is the subject of discussion, the finished print or on-screen image more commonly lends a photograph its macro status. For example, when producing a 6×4-inch (15×10-cm) print using 35 format (36×24 mm) film or sensor, a life-size result is possible with a lens having only a 1:4 reproduction ratio. Reproduction ratios much greater than 10:1 are considered to be photomicrography, often achieved with digital microscope (photomicrography should not be confused with microphotography, the art of making very small photographs, such as for microforms). Due to advances in sensor technology, today's small-sensor digital cameras can rival the macro capabilities of a DSLR with a "true" macro lens, despite having a lower reproduction ratio, making macro photography more widely accessible at a lower cost. In the digital age, a "true" macro photograph can be more practically defined as a photograph with a vertical subject height of 24 mm or less. History The term photo-macrograph was proposed in 1899 by W. H. Walmsley for close-up images with less than 10 diameters magnification, to distinguish from true photo-micrographs. Development of the photo-micrograph led to the evolution of macro photography. One of the earliest pioneers of macro photography was Percy Smith, born in 1880. He was a British nature documentary filmmaker, and was known for his close-up photographs. Equipment and techniques "Macro" lenses specifically designed for close-up work, with a long barrel for close