populations took place when Albanians dwelt in forests 600–900 metres above sea level. Rusakov notes that almost all lexemes related to seamanship in Albanian are loan-words, which may indicate that speakers of the proto-language did not live on the Adriatic coast or in close proximity to it. Greek Linguistic contact between Albanian and Greek has been securely dated to the Iron Age. Previous contacts may include contacts between the respective post-PIE languages which gave rise to the two languages. According to Huld (1986), Ancient Greek loans originate from two distinct geographical groups; Greek-speaking populations from ancient Macedonia and the Greek colonies on the Adriatic coast. It has been known since the 1910 work of German linguist Albert Thumb, that Albanian possesses a small number of Greek loans as old or older than its earliest Latin loans. Krzysztof (2016) suggests that Doric Greek may have also received some loanwords from Proto-Albanian, that could be as early as the 7th century BC; he specifically points to seven words recorded by the Greek grammarian Hesychius of Alexandria (5th century AD), and particularly to the term 'a kind of earring', which was first attested in the work of the choral lyric poet Alcman. Words borrowed from Greek (e.g. Gk (NW) mākhaná "device, instrument" > mokër "millstone", Gk (NW) drápanon > drapër "sickle" etc.) date back before the Christian era and are mostly of the Doric Greek dialect, which means that the ancestors of the Albanians were in contact with the northwestern part of Ancient Greek civilization and probably borrowed words from Greek cities (Dyrrachium, Apollonia, etc.) in the Illyrian territory, colonies which belonged to the Doric division of Greek, or from contacts in the Epirus area. The earliest Greek loans began to enter Albanian circa 600 BC, and are of Doric provenance, tending to refer to vegetables, fruits, spices, animals and tools. Joseph argues that this stratum reflects contacts between Greeks and Proto-Albanians from the 8th century BC onward, with the Greeks being either colonists on the Adriatic coast or Greek merchants inland in the Balkans. The second wave of Greek loans began after the split of the Roman empire in 395 and continued throughout the Byzantine, Ottoman and modern periods. An argument in favor of a northern origin for Albanian is the relatively small number of loanwords from Ancient Greek, mostly from Doric dialect, even though Southern Illyria neighbored the Classical Greek civilization