from the western lineage evolved the Bubal and western hartebeest. The southern lineage gave rise to Lichtenstein's and red hartebeest. These two taxa are phylogenetically close, having diverged only 0.2 million years ago. The study concluded that these major events throughout the hartebeest's evolution are strongly related to climatic factors, and that there had been successive bursts of radiation from a more permanent population—a refugium—in eastern Africa; this could be vital to understanding the evolutionary history of not only the hartebeest but also other mammals of the African savanna. The earliest fossil record dates back to nearly 0.7 million years ago. Fossils of the red hartebeest have been found in Elandsfontein, Cornelia (Free State) and Florisbad in South Africa, as well as in Kabwe in Zambia. In Israel, hartebeest remains have been found in northern Negev, Shephelah, Sharon Plain and Tel Lachish. This population of the hartebeest was originally limited to the open country of the southernmost regions of the southern Levant. It was probably hunted in Egypt, which affected the numbers in the Levant, and disconnected it from its main population in Africa. Description A large antelope with a particularly elongated forehead and oddly shaped horns, the hartebeest stands just over at the shoulder, and has a typical head-and-body length of . The weight ranges from . The tail, long, ends in a black tuft. The other distinctive features of the hartebeest are its long legs (often with black markings), short neck, and pointed ears. A study correlated the size of hartebeest species to habitat productivity and rainfall. The western hartebeest is the largest subspecies, and has a characteristic white line between the eyes. The red hartebeest is also large, with a black forehead and a contrasting light band between the eyes. The large Lelwel hartebeest has dark stripes on the front of its legs. Coke's hartebeest is moderately large, with a shorter forehead and longer tail in comparison to the other subspecies. Lichtenstein's hartebeest is smaller, with dark stripes on the front of the legs, as in the Lelwel hartebeest. The Swayne's hartebeest is smaller than the Tora hartebeest, but both have a shorter forehead and similar appearance. Generally short and shiny, the coat varies in colour according to subspecies. The western hartebeest is a pale sandy-brown, but the front of the legs are darker. The red hartebeest is a reddish-brown, with a dark face. Black markings can