The Greco-Buddhist art or Gandhara art of the north Indian subcontinent is the artistic manifestation of Greco-Buddhism, a cultural syncretism between Ancient Greek art and Buddhism. It had mainly evolved in the ancient region of Gandhara. The series of interactions leading to Gandhara art occurred over time, beginning with Alexander the Great's brief incursion into the area, followed by the Mauryan Emperor Ashoka converting the region to Buddhism. Buddhism became the prominent religion in the Indo-Greek Kingdoms. However, Greco-Buddhist art truly flowered and spread under the Kushan Empire, when the first surviving devotional images of the Buddha were created during the 1st-3rd centuries CE. Gandhara art reached its zenith from the 3rd-5th century CE, when most surviving motifs and artworks were produced. Gandhara art is characterized by Buddhist subject matter, sometimes adapting Greco-Roman elements, rendered in a style and forms that are heavily influenced by Greco-Roman art. It has the strong idealistic realism and sensuous description of Hellenistic art, and it is believed to have produced the first representations of Gautama Buddha in human form, ending the early period of aniconism in Buddhism. The representation of the human form in large sculpture had a considerable influence, both to the south in the rest of India, and to the east, where the spread of Buddhism carried its influence as far as Japan. Historical outline There has been disagreement among art historians as to whether Gandharan art owes more to the culture established immediately after Alexander's campaigns, or to a synthesis several centuries later between travelling Greco-Roman artists from the eastern Roman Empire in regions where Greek settlers were still important. Most of the considerable quantity of Gandharan art that can be dated comes from after about 50 AD, although some clearly was created earlier. For this reason, some scholars prefer to call this Romano-Indian art, or talk of an "Indo-Classical style". The French scholar Alfred Foucher first identified the Western influences on Gandharan art at the end of the 19th century. He was initially a proponent of the continuity between the first Greek settlements and this art, and dated much of the art much earlier than more recent scholars do. However, he later revised his views and datings somewhat. His views as to dates and the crucial period of Western influence came to be widely rejected, but then received considerable support by the discovery of the important deserted city site of