scholars, such as Anderson (1929), and later Gill (1951, 1952, 1954), thereby leaving the issue unresolved. In 1981, another paper was published arguing that certain cuts to bones of large marsupials had been caused by Thylacoleo. This paper by Horton and Wright was able to counter earlier arguments that such marks were the result of humans, largely by pointing out the presence of similar marks on the opposite side of many bones. They concluded that humans were extremely unlikely to have made the marks in question, but that if so "they had set out to produce only marks consistent with what Thylacoleo would produce". Since then, the academic consensus has emerged that Thylacoleo was a predator and a hypercarnivore. Besides the most common hypothesis that it was an active predator, a variety of other theories existed in the late 19th to early 20th centuries as to the diet and feeding of Thylacoleo, with hypotheses of it being a scavenger filling the ecological niche of hyenas, being a specialist of crocodile eggs, or even a melon-eater. As late as 1954, doubts were still being raised as to whether it was actually a hypercarnivore. The marsupial lion's limb proportions and muscle mass distribution indicate that, although it was a powerful animal, it was not a particularly fast runner. Paleontologists conjecture that it was an ambush predator, either sneaking up and then leaping upon its prey, or dropping down on it from overhanging tree branches. This is consistent with the depictions of the animal as striped: camouflage of that kind is needed for stalking and hiding in a largely forested habitat (like tigers) rather than chasing across open spaces (like lions). Trace fossils in the form of claw marks and bones from caves in Western Australia analyzed by Gavin Prideaux et al. indicate marsupial lions could also climb rock faces, and likely reared their young in such caves as a way of protecting them from potential predators. It is thought to have hunted large animals such as the enormous Diprotodon and giant browsing kangaroos like Sthenurus and Procoptodon, and competed with other predatory animals such as the giant monitor lizard, Megalania, and terrestrial crocodiles such as Quinkana. The marsupial lion may have cached kills in trees in a manner similar to the modern leopard. Like many predators, it was probably also an opportunistic scavenger, feeding on carrion and driving off less powerful predators