their prime environment appears to be steep ravine-indented slopes along artificial watercourses, with the occasional large dead laurel tree and much tree heath. This species is highly mobile between different areas at different times of year. It was very abundant when the islands were first colonised by humans, but was extirpated. The losses on the islands were largely due to deforestation for wood and to create agricultural and grazing land. The exclusion of livestock from the native forest allows it to regenerate and create more suitable habitat. Some illegal hunting and poisoning continues because of the damage this pigeon can do to crops. Perhaps the main limiting factor on the rate at which the pigeon increases its numbers is eggs and young being taken by introduced black rats. It is now classed as Near Threatened on the IUCN Red List, an improvement on its Threatened status in 1988. This species is protected under the European Union Birds Directive, and the laurel forests under the Habitats Directive. Taxonomy The genus Columba is the largest within the pigeon family, and has the widest distribution. Its members are typically pale grey or brown, often with white head or neck markings or iridescent green or purple patches on the neck and breast. The neck feathers may be stiffened and aligned to form grooves. One of several subgroups within Columba consists of the widespread Eurasian common wood pigeon, Bolle's pigeon, the Trocaz pigeon, and the African Afep pigeon. The three Macaronesian endemic pigeons, this species, Bolle's and Trocaz, are thought to be derived from isolated island populations of C. palumbus. The Atlantic archipelagos of the Canaries, Azores, and Madeira have a volcanic origin and they have never been part of a continent. The formation of Madeira started in the Miocene and the island was substantially complete by 700,000 years ago. At various times in the past, the major islands of these archipelagos were all colonised by ancestral wood pigeons, which evolved on their respective islands in isolation from the mainland populations. Mitochondrial and nuclear DNA sequences suggest that the ancestor of Bolle's pigeon may have arrived in the Canaries about 5 mya, but an older lineage that gave rise to another Canarian endemic, the laurel pigeon, C. junoniae, may date from 20 mya. The most recent wood pigeon arrival on Macaronesia was that which gave rise to the subspecies . The Trocaz pigeon was formally