is under anthropogenic pressures that include infrastructure development and pollution. Less than two percent of the area is under federal protection. Researchers have warned that the cumulative effect of various minor stressors could lead to a more catastrophic collapse if usage of the Pantanal is not more strictly regulated. References Fish described in 1891 Taxa named by Eduardo Ladislao Holmberg Characiformes Fish of South America Fish of Paraguay Fish of Argentina Fish of Brazil