recognisable in some fossil crustaceans; and entoniscidae living inside some species of crab and shrimp. Cymothoa exigua is a parasite of the spotted rose snapper Lutjanus guttatus in the Gulf of California; it causes the tongue of the fish to atrophy and takes its place in what is believed to be the first instance discovered of a parasite functionally replacing a host structure in animals. Reproduction and development In most species, the sexes are separate and there is little sexual dimorphism, but a few species are hermaphroditic and some parasitic forms show large differences between the sexes. Some Cymothoidans are protandrous hermaphrodites, starting life as males and later changing sex, and some Anthuroideans are the reverse, being protogynous hermaphrodites that are born female. Some Gnathiidans males are sessile and live with a group of females. Males have a pair of penises, which may be fused in some species. The sperm is transferred to the female by the modified second pleopod which receives it from the penis and which is then inserted into a female gonopore. The sperm is stored in a special receptacle, a swelling on the oviduct close to the gonopore. Fertilisation only takes place when the eggs are shed soon after a moult, at which time a connection is established between the semen receptacle and the oviduct. The eggs, which may number up to several hundred, are brooded by the female in the marsupium, a chamber formed by flat plates known as oostegites under the thorax. This is filled with water even in terrestrial species. The eggs hatch as mancae, a post-larval stage which resembles the adult except for the absence of the last pair of pereopods. The lack of a swimming phase in the life cycle is a limiting factor in isopod dispersal, and may be responsible for the high levels of endemism in the order. As adults, isopods differ from other crustaceans in that moulting occurs in two stages known as "biphasic moulting". First they shed the exoskeleton from the posterior part of their body and later shed the anterior part. The giant Antarctic isopod Glyptonotus antarcticus is an exception, and moults in a single process. Terrestrial isopods The majority of crustaceans are aquatic and the isopods are one of the few groups of which some members now live on land. The only other crustaceans which include a small number of terrestrial species are amphipods (like