taxa of different sizes, until the discovery of Kelenken. Evolution In their 2007 description, Bertelli and colleagues classified Kelenken as a member of the family Phorusrhacidae, based on its enormous size, combined with its sideways compressed, strongly hooked beak (or rostrum, the part of the jaws that formed the beak), and convex culmen (the top of the upper beak). Five phorusrhacid subfamilies were recognized at the time (Brontornithinae, Phorusrhacinae, Patagornithinae, Mesembriornithinae, and Psilopterinae), though their validity had not then been confirmed through cladistic analysis, and the describers found Kelenken most similar to taxa that had traditionally been considered phorusrhacines. Features shared with phorusrhacines include that the hind part of the skull is low and compressed from top to bottom, a wide occipital table, a blunt postorbital process, and a tarsometatarsus that is similar to that of Titanis in that the supratrochlear surface of the lower end is flat. Further comparison was hampered by the lack of anatomical information about phorusrhacines. The Brazilian paleontologist Herculano Alvarenga and colleagues published a phylogenetic analysis of Phorusrhacidae in 2011 that found Kelenken and Devincenzia to be sister taxa, each other's closest relatives. While the analysis supported there being five subfamilies, the resulting cladogram did not separate Brontornithinae, Phorusrhacinae, and Patagornithinae. In their 2015 description of Llallawavis, the Argentinian paleontologist Federico J. Degrange and colleagues performed a phylogenetic analysis of Phorusrhacidae, wherein they found Phorusrhacinae to be polyphyletic (an unnatural grouping). The following cladogram shows the position of Kelenken following the 2015 analysis: During the early Cenozoic, after the extinction of the non-bird dinosaurs, mammals underwent an evolutionary diversification, and some bird groups around the world developed a tendency towards gigantism; this included the Gastornithidae, the Dromornithidae, the Palaeognathae, and the Phorusrhacidae. Phorusrhacids are an extinct group within Cariamiformes, the only living members of which are the two species of seriemas in the family Cariamidae. While they are the most speciose group within Cariamiformes, the interrelationships between phorusrhacids are unclear due to the incompleteness of their remains. Phorusrhacids were present in South America from the Paleocene (when the continent was an isolated island) and survived until the Pleistocene. They also appeared in North America at the end of the Pliocene, during the Great American Biotic Interchange, and while fossils from Europe have been assigned to the group, their classification is disputed. It is unclear where the group originated; both cariamids and phorusrhacids may have arisen