Juniperus macrocarpa (large-fruited juniper, syn. J. oxycedrus subsp. macrocarpa (Sibth. & Sm.) Ball) is a species of juniper, native across the northern Mediterranean region from southwestern Spain east to western Turkey and Cyprus, growing on coastal sand dunes from sea level up to in altitude. It is a spreading shrub tall, rarely a small tree up to tall. The leaves are broad lanceolate, produced in whorls of three, green, long and 2–3 mm broad, with a double white stomatal band split by a green midrib on the inner surface. It is dioecious, with separate male and female plants. The seed cones are berry-like, green ripening in 18 months to orange-red with a variable pink waxy coating; they are spherical, 12–18 mm diameter, and have six fused scales in two whorls, three of the scales with a single seed. The seeds are dispersed when birds eat the cones, digesting the fleshy scales and passing the hard seeds in their droppings. The pollen cones are yellow, 2–3 mm long, and fall soon after shedding their pollen in late winter. Despite its distinct morphology with large cones and broad leaves more like those of Juniperus drupacea, it has often been treated as a subspecies of Juniperus oxycedrus, though recent genetic studies have shown its DNA is distinct from that of J. oxycedrus. References macrocarpa Trees of Mediterranean climate Trees of Europe Trees of Turkey Flora of Cyprus Dioecious plants Flora of the Mediterranean Basin