). Other archaic human species are assumed to have spread throughout Africa by this time, although the fossil record is sparse. Their presence is assumed based on traces of admixture with modern humans found in the genome of African populations. Homo naledi, discovered in South Africa in 2013 and tentatively dated to about 300,000 years ago, may represent fossil evidence of such an archaic human species. Neanderthals spread across the Near East and Europe, while Denisovans appear to have spread across Central and East Asia and to Southeast Asia and Oceania. There is evidence that Denisovans interbred with Neanderthals in Central Asia where their habitats overlapped. Neanderthal evidence has also been found quite late at 33,000 years ago at the 65th latitude of the Byzovaya site in the Ural Mountains. This is far outside of any otherwise known habitat, during a high ice cover period, and perhaps reflects a refugia of near extinction. Homo sapiens Dispersal throughout Africa Homo sapiens are believed to have emerged in Africa about 300,000 years ago, based in part on thermoluminescence dating of artifacts and remains from Jebel Irhoud, Morocco, published in 2017. The Florisbad Skull from Florisbad, South Africa, dated to about 259,000 years ago, has also been classified as early Homo sapiens. Previously, the Omo remains, excavated between 1967 and 1974 in Omo National Park, Ethiopia, and dated to 200,000 years ago, were long held to be the oldest known fossils of Homo sapiens. In September 2019, scientists reported the computerized determination, based on 260 CT scans, of a virtual skull shape of the last common human ancestor to anatomically modern humans, representative of the earliest modern humans, and suggested that modern humans arose between 260,000 and 350,000 years ago through a merging of populations in East and South Africa. In July 2019, anthropologists reported the discovery of 210,000 year old remains of a H. sapiens and 170,000 year old remains of a H. neanderthalensis in Apidima Cave in southern Greece, more than 150,000 years older than previous H. sapiens finds in Europe. Early modern humans expanded to Western Eurasia and Central, Western and Southern Africa from the time of their emergence. While early expansions to Eurasia appear not to have persisted, expansions to Southern and Central Africa resulted in the deepest temporal divergence in living human populations. Early modern human expansion in sub-Saharan Africa appears to have contributed to the end of