a unique species, most notably its unique snout, which does not strongly taper towards the tip as in female Pteranodon specimens, with the upper and lower margins running almost parallel instead. Kellner thought this meant the specimen must have had a crest running along the length of the snout. However, a re-examination of the fossil published in 2017 by Elizabeth Martin-Silverstone and colleagues argued that the differences in the snout were more likely due to male pteranodontids having longer, broader bills than the more complete female specimens Kellner used for comparison. Martin-Silverstone concluded that "Dawndraco" was simply a male G. sternbergi with a very long bill with a gradual taper. Geosternbergia fossils are known from the Niobrara and Sharon Springs Formations of the central United States. Geosternbergia existed as a group for more than four million years during the late Coniacian – early Campanian stages of the Cretaceous period. The genus is present in the lower the Niobrarra Formation except for the upper two; in 2003, Kenneth Carpenter surveyed the distribution and dating of fossils in this formation, demonstrating that Geosternbergia sternbergi existed there from 88–85 million years ago, while the species later named G. maiseyi existed between 81.5 and 80.5 million years ago. Description Size Geosternbergia was among the largest pterosaurs, with the wingspan of most adults ranging between . No complete skulls of adult males have been found, but a nearly complete lower jaw has been estimated at long. While most specimens are found crushed, enough fossils exist to put together a detailed description of the animal. Geosternbergia sternbergi was very similar to the more well-known pterosaur species Pteranodon longiceps and is often considered simply an earlier species of Pteranodon itself. It is different from Pteranodon mainly due to its earlier time period and broader, more upright crest. Skull and beak The upright cranial crest of Geosternbergia is its distinctive characteristic. These crests consisted of skull bones (frontals) projecting upward and backward from the skull. The size and shape of these crests varied due to a number of factors, including age, sex, and species. Male G. sternbergi, the older species of the two described to date, had a larger vertical crest with a broad forward projection, while G. maiseyi had a short, rounded vertical crest and was generally smaller. Females of both species were smaller and bore small, rounded crests. The crests were probably mainly display structures, though