is a miniaturised species, attaining an adult body size of 11–12 mm. It was argued to be a proportional dwarf in its original description, based on its non-paedomorphic proportions compared to similarly sized Cophylinae frogs. Reproductive Ecology The mating habits of most species of Rhombophryne are not known. It is likely that all members of this genus engage in parental care, which is common to most members of the Cophylinae. Two adult R. testudo individuals were found together with 18 putative offspring in a burrow by a stream at the edge of Lokobe Reserve on Nosy Be, at the end of the dry season. This is the only direct evidence of reproductive behaviour in this genus. So far, the advertisement calls of eight species have been published: R. botabota, R. minuta, R. mangabensis, R. matavy, R. testudo, R. nilevina, R. coronata, and R. proportionalis. Most of their calls are honking sounds emitted after quite long intervals, but R. minuta emits paired notes, and R. proportionalis emits series of 9–17 notes. Species Rhombophryne botabota Scherz, Glaw, Vences, Andreone & Crottini, 2016 Rhombophryne coronata (Vences and Glaw, 2003) Rhombophryne coudreaui (Angel, 1938) Rhombophryne diadema Scherz, Hawlitschek, Andreone, Rakotoarison, Vences & Glaw, 2017 Rhombophryne guentherpetersi (Guibé, 1974) Rhombophryne laevipes (Mocquard, 1895) Rhombophryne longicrus Scherz, Rakotoarison, Hawlitschek, Vences & Glaw, 2015 Rhombophryne mangabensis Glaw, Köhler & Vences, 2010 Rhombophryne matavy D'Cruze, Köhler, Vences & Glaw, 2010 Rhombophryne minuta (Guibé, 1975) Rhombophryne nilevina Lambert, Hutter & Scherz, 2017 Rhombophryne ornata Scherz, Ruthensteiner, Vieites, Vences & Glaw, 2015 Rhombophryne proportinalis Scherz, Hutter, Rakotoarison, Riemann, Rödel, Ndriantsoa, Glos, Roberts, Crottini, Vences & Glaw, 2019 Rhombophryne regalis Scherz, Hawlitschek, Andreone, Rakotoarison, Vences & Glaw, 2017 Rhombophryne savaka Scherz, Glaw, Vences, Andreone & Crottini, 2016 Rhombophryne serratopalpebrosa (Guibé, 1975) Rhombophryne tany Scherz, Ruthensteiner, Vieites, Vences & Glaw, 2015 Rhombophryne testudo Boettger, 1880 Rhombophryne vaventy Scherz, Ruthensteiner, Vences & Glaw, 2014 Rhombophryne ellae Scherz, 2020 References Endemic frogs of Madagascar Amphibian genera Taxa named by Oskar Boettger Taxonomy articles created by Polbot