According to the IUCN it was threatened by habitat loss in 2004. Protected populations occur in Carara National Park, Río Piro Wildlife Refuge, Corcovado National Park, Maquenque National Wildlife Refuge (from ), and at least historically in the Monteverde Cloud Forest Reserve area. References External links Numerous in situ photographs of the frog rugosus Amphibians described in 1873 Taxa named by Wilhelm Peters Taxonomy articles created by Polbot