Boophis is the only genus in the mantellid frog subfamily Boophinae. They are commonly known as bright-eyed or skeleton frogs. They show typical 'tree frog' traits, and are a good example of convergent evolution with morphologically similar species in the families Hylidae and Rhacophoridae, among others. This genus can only be found on Madagascar and Mayotte Island (Comoros). Taxonomy The genus Boophis was described by Johann Jakob von Tschudi in 1838. It was originally considered a member of the African-Asian family Rhacophoridae, but was moved to the family Mantellidae in 2001, into its own subfamily, Boophinae Vences & Glaw, 2001. Morphology Boophis are arboreal frogs, exhibiting many traits that define 'tree frogs', such as expanded toe discs, long hindlimbs, and large eyes. Boophis are especially characterised by bright colouration of the iris, which is typically intricately patterned towards its inside, and often green or blue, but occasionally also red, purple, or yellow in the outer iris area. This has led to the vernacular name of bright-eyed frogs for the genus. Many species of Boophis have almost translucent skin, allowing bones and internal organs to be observed as in the unrelated glass frogs (Centrolenidae) of the tropical Americas. This has led to the vernacular name skeleton frogs for some members of the genus. Species The genus has presently nearly 80 species; new ones are being described every few months on average. This list may not be exhaustive. Boophis albilabris (Boulenger, 1888) Boophis albipunctatus Glaw & Thiesmeier, 1993 Boophis andohahela Andreone, Nincheri & Piazza, 1995 Boophis andrangoloaka (Ahl, 1928) Boophis andreonei Glaw & Vences, 1994 Boophis anjanaharibeensis Andreone, 1996 Boophis ankarafensis Penny, Andreone, Crottini, Holderied, Rakotozafy, Schwitzer & Rosa, 2014 Boophis ankaratra Andreone, 1993 Boophis arcanus Glaw, Köhler, de la Riva, Vieites & Vences, 2010 Boophis axelmeyeri Vences, Andreone & Veities, 2005 Boophis baetkei Köhler, Glaw & Vences, 2008 Boophis blommersae Glaw & Vences, 1994 Boophis boehmei Glaw & Vences, 1992 Boophis boppa Hutter & Lambert, 2015 Boophis bottae Vences & Glaw, 2002 Boophis brachychir Boettger, 1882 Boophis burgeri Glaw & Vences, 1994 Boophis calcaratus Vallan, Vences & Glaw, 2010 Boophis doulioti Angel, 1934 Boophis elenae Andreone, 1993 Boophis englaenderi Glaw & Vences, 1994 Boophis entingae Glaw, Köhler, de la Riva, Vieites & Vences, 2010 Boophis erythrodactylus (Guibé, 1953) Boophis fayi Köhler, Glaw, Rosa, Gehring, Pabijan, Andreone & Vences, 2011 Boophis feonnyala Glaw, Vences, Andreone & Vallan, 2001 Boophis goudotii Tschudi, 1838