he did not formally synonymise this taxon as well. In 1921 Emmett Dunn collected what was thought to be the first specimen of E. rugosus known from Costa Rica near the town of Monteverde, himself and Tom Barbour not agreeing with Noble's synonymy. Fritz Nieden also ignored Noble in his contribution to the second edition of Das Tierreich (book 46) in 1923. Between 1947 and 1951 Richard Clark Taylor and Edward Harrison Taylor visited Costa Rica for a combined total of 8 months, and the two men amassed a collection of over 6,000 herpetology specimens, more than the total that had been harvested from the country in the previous centuries. In his report on the amphibians of the country based on this collection, the latter Taylor follows Noble in considering E. megacephalus to be a synonym E. rugosus, however Taylor expresses some doubt as to if E. pelviculus and E. gulosus could truly be synonymised with E. rugosus, both based on such a paucity of unclear evidence. Throughout most of 20th century this species was considered to be two accepted species under Stejneger's Eleutherodactylus taxonomy: E. florulentus (still only known from a single damaged specimen, missing since the 19th century) and E. rugosus (at the time a mix of both modern E. rugosus and modern E. megacephalus, and thought to be distributed from Honduras (perhaps even Mexico) to Panama), but in 1975 John D. Lynch synonymised both E. florulentus and E. rugosus, along with the older nomen E. pelviculus and E. megacephalus, and the mysterious E. gulosus (also still only known from the initial collections at the time), with E. biporcatus -under which the frogs of this species were known for the next few decades, and which was henceforth thought to be distributed from Honduras to Peru. In 2002 Savage and Myers reassessed E. biporcatus, searching for the type specimens of the various synonyms in Berlin. They were unable to locate some (see above), but managed to discover that the actual holotype for E. biporcatus had been collected in Venezuela, not in Veraguas Province as had been thought previously. This meant that E. maussi, as a Venezuelan taxon of frogs was known at the time, was in fact a junior synonym of E. biporcatus. Due to this, Savage and Myers split the old E. biporcatus into three resurrected taxa and one new species (E. opimus). Savage and Myers also synonymised