Yaeyama Islands, as the species certainly does not occur on Okinawa. And thus, the frogs formerly known as R. psaltes, which had also been found on Taiwan in the meantime, turned out to be the same species that Boettger had described, and hence according to ICZN rules is now known as R. okinavana. Footnotes References (1895a): Neue Frösche und Schlangen von den Liukiu-Inseln ["New frogs and snakes from the Ryūkyū Islands"]. [In German] Bericht des Offenbacher Verein für Naturkunde 33-36: 101–117. (1895b): Neue Frösche und Schlangen von den Liukiu-Inseln ["New frogs and snakes from the Ryūkyū Islands"]. [In German] Zoologischer Anzeiger 18: 266–270. (1985): A new frog (genus Rana) from the Yaeyama group of the Ryukyu Islands. Herpetologica 41: 150–158. (2007): Unmasking Rana okinavana Boettger, 1895 from the Ryukyus, Japan (Amphibia: Anura: Ranidae). Zool. Sci. 24: 199–204. (HTML abstract) (1907): Herpetology of Japan and adjacent territory. Bulletin of the United States National Museum 58 1–577. External links Mating Ryūkyū brown frogs photo. Retrieved 2008-FEB-02. Kampira Falls frog photo. Retrieved 2008-FEB-02. Rana (genus) Amphibians of Japan Taxonomy articles created by Polbot