neo-tropical and considered the smallest urodeles in the world. The genus Desmognathus consists of nineteen species which all vary in body size, life history, and habitat. The most basic way of sorting the species in this genus is by body size and distance they are found from streams which they inhabit. In 2010, there was a taxonomic revision to Desmognathus wrighti. This revision was based on genetic and ecological analyses. This divergence has been reported by DNA sequencing, but further research shows several differences in the body type and ventral pigmentation. An ecological niche analysis was completed on two niches separated by the French Broad River and significant morphological variation was found between the two lineages. Findings show differences in snout-vent length, body condition, and head width, suggesting that there are two distinct and separately evolving lineages in Desmognathus wrighti. The diverging northern lineage, Desmognathus organi, is the new species described by Crespi and Browne in 2010. The revision to the taxonomic Desmognathus wrighti was seen needed for the northern pygmy salamander to reflect the genetic, ecological, and geographical differences in the two different populations located north and south of the French Broad River. The original species Desmognathus wrighti described by King more accurately describes the lineage from the southern part of the French Broad River and the morphological descriptions described on this page. The new species Desmognathus organi, the Northern pygmy salamander, is a sister taxon to D. wrighti. Differences of D. organi from its sister taxon includes a slightly larger body length, wider heads, and the ventral gold iridophore patterns are absent. Also, D. organi is found in higher elevations north of the French Broad River including Tennessee, North Carolina, and Virginia. Geographical distribution Desmognathus wrighti has a very fragmented geographical distribution. The species is found in the Southern Appalachian Mountains of North Carolina and Tennessee south of the French Broad River. In Tennessee, D. wrighti inhabits the Great Smoky Mountains National Park in Blount, Cocke, and Sevier counties. The species is also found in the Cherokee National Forest in Monroe County. In North Carolina, the pygmy salamander is only documented in Graham County along the Tennessee border. The highest abundance of pygmy salamanders is found at higher elevations ranging from 1600 to 2082 meters. The distribution of the salamander primarily lies adjacent to areas of red spruce (Picea rubens) and Fraser's fir (Abies fraseri) forests at these