been found in bromeliads. The eggs are laid on dry land. This species possesses "direct development", the eggs develop directly into tiny juvenile frogs, skipping a tadpole stage. Diseases This frog is suspected to be susceptible to the deadly pathogenic fungus Batrachochytrium dendrobatidis. Conservation It was formerly believed to be endemic to Costa Rica. It was assessed for the IUCN as being "critically endangered" in 2004, and in 2008 it retained this status, due to its "drastic population decline". It is considered threatened in Costa Rica, and is protected by the wildlife conservation law No. 7317, environmental law No. 7554 and the decree No. 26435-MINAE. The frog was regularly seen in the annual collections of amphibians at least in two of the various nature parks of Costa Rica it was known from, but in the late 1980s this species suddenly declined and disappeared from the amphibian collections. The last record of this species was from 1990, and it has not been seen since (as of August 2007), despite it having been searched for. It has not been seen in Cascajal, San Jose Province, since 1972. Because it has seemingly disappeared from the areas in which it has almost only been collected - otherwise pristine, well-managed nature reserves, infection by the introduced fungal disease chytridiomycosis has been suggested as the reason of decline, as this has also impacted other similar frogs from similar habitats. The IUCN also mentions unspecified possible impacts from climate change. It is, or was, found in the Monteverde Cloud Forest Reserve, the Alberto Manuel Brenes Biological Reserve and the Maquenque National Wildlife Refuge in Costa Rica from in altitude. There is no ex situ population of this species under conservation. References External links Undated photograph of an individual from Bocas del Toro, Panama. andi Amphibians of Costa Rica Amphibians of Panama Amphibians described in 1974 Taxa named by Jay M. Savage Taxonomy articles created by Polbot