Theodore Galloway: A national security adviser who often chastises NEST and the Autobots for their destructive tactics. Glenn Morshower as General Morshower: The supervisor of NEST. Matthew Marsden as Graham: a British Army NEST officer. Rainn Wilson as Professor R.A. Colan: Sam and Leo's astronomy teacher. Michael Papajohn as Colin "Cal" Banes: Mikaela's father. Voices Peter Cullen as Optimus Prime: The leader of the Autobots who transforms into a blue and red 1994 Peterbilt 379 semi-trailer truck. Hugo Weaving as Megatron: The Fallen's apprentice and the leader of the Decepticons who transforms into a Cybertronian tank capable of flight. Tony Todd as The Fallen: A Prime who is the first and founder of the Decepticons, as well as the master of Megatron. Mark Ryan as Jetfire: A former Decepticon Seeker turned Autobot who transforms into a SR-71 Blackbird. Jess Harnell as Ironhide: The Autobot weapons specialist and Optimus's new second-in-command who transforms into a black 2009 GMC Topkick C4500. Robert Foxworth as Ratchet: The Autobot medical officer who transforms into a yellow 2009 search and rescue Hummer H2 ambulance. Charlie Adler as Starscream: Megatron's second-in-command who transforms into a Lockheed Martin F-22 Raptor. Frank Welker as: Soundwave: The Decepticon communications officer who transforms into a Cybertronian satellite. He is only seen orbiting Earth in this film. Reedman: A razor-thin Decepticon composed of thousands of miniature bead-like Decepticons. Devastator: A massive Decepticon who is the combination of nine construction vehicles. In the Transformers lore, these vehicles are meant to be individual Decepticons called the Constructicons. Concept art was even created for their robot forms to be featured in the film, but most of them (except for Demolishor and Rampage) ended up not being used, only being featured in the film's related toy line.The vehicles that compose Devastator are: A black and silver Mack concrete mixer truck (head of Devastator, originally a Constructicon called Mixmaster) A yellow Caterpillar 992G scoop loader (right arm, originally a Constructicon called Scrapper) A yellow Volvo EC700C crawler excavator fitted with a Stanley UP 45SV attachment (originally a Constructicon called Scrapmetal) A green Caterpillar 773B dump truck (right leg, originally a Constructicon called Long Haul) A red and white Terex O&K RH 400 excavator, like Demolishor (torso, originally a Constructicon called Scavenger) A yellow Kobelco CKE2500 II crawler crane (left arm, originally a Constructicon called Hightower) A red KW Dart D4661 Tractor Truck articulated dump truck (lower