Romulus is a fictional supervillain appearing in American comic books published by Marvel Comics. The character is featured in particular in those titles featuring Wolverine. He is the leader of the Lupines, a species resembling humans that he claims evolved from canines instead of primates through convergent evolution. A shadowy character whose origin and motives remain a mystery, he is shown to have orchestrated most major events in the life of Wolverine, manipulating and controlling him for most of his life. Publication history Created by writer Jeph Loeb and artist Simone Bianchi, the character first appeared obscured by shadow in Wolverine (vol. 3) #50 (March 2007), and made his first full appearance in Wolverine: Origins #39 (August 2009). Fictional character biography Romulus is first seen in several flashbacks that Wolverine experiences, later explained by Wild Child to have been induced by Romulus himself. He is shown leading the Lupines in prehistoric and barbaric times, being an emperor in Ancient Rome, as well as being the main force behind Weapon X. In the present, Romulus restored Feral and Thornn's feline look after their depowerment on M-Day, augmented Wild Child's powers greatly, and seemed to have erased all humanity from Sabretooth causing the latter's death at the hands of Wolverine (later it was revealed Sabretooth was a clone). Romulus did all of this to reveal himself to Wolverine. Romulus was also the one who took Daken from his dead mother's womb and raised him as a ruthless killer. Nick Fury revealed to Wolverine that Romulus has been manipulating people from his family's bloodline for centuries, and that he plans for Daken to become the next-generation Weapon X. This was achieved through Tinkerer who bonded the metal from the Muramasa blade to Daken's inner claws. Daken hinted he complied with the process because of his own agenda. Romulus told Daken to kill all the Weapon X experiments that gave them a healing factor with his Muramasa claws. He killed every one except Deadpool. He appears to have ties with Russian authorities, arranging Omega Red's transfer to a regular Russian prison to set a trap for Wolverine. After Wolverine temporarily subdues Omega Red, he is captured by Wild Child, who reveals that Romulus made Logan and several villains battle each other, his agenda being vaguely described as him needing a successor. Moments after Wolverine uses the Muramasa blade to kill Omega Red, Romulus reveals