first finger. The toes and the outer two fingers of V. ritae are webbed; the first two fingers (of which the first is a bit longer) are completely free. The webbing formula for the outer fingers is III (2−-21/3) – (1+-2−) IV; for the toes, it is I 1 – (2−-2) II (1-1+) – (2-21/4) III (1+-11/2) – 2+ IV (2-21/3) – 1 V. The disc at the tip of the third finger is mid-sized, larger than the eardrum but less than half the diameter of the eye. This species has tubercles on the thighs below the vent, but only low folds with no iridophores on the ulna and inner tarsus. Ecology Its natural habitats are tropical moist lowland forests and rivers; it is usually found in riparian vegetation. Occurring at elevations below ASL; it is apparently also absent from the coastlands up to some dozen meters above sea level. The calls of this species have supposedly been described at least twice, but it is not clear whether they are actually of V. ritae; one description appears to be of C. midas calls instead. The other notes that the males call sitting on the upperside of leaves, usually giving very brief (0.10–0.15 seconds) calls that are loudest around 4640–5160 Hz singly or doubly, but apparently never three in a row – Helena's glass frog (V. helenae) gives double or triple but apparently never single calls. It is unknown if and how the males physically fight for females. The clutches are deposited on either side of leaves above small streams; after hatching the tadpoles drop into the water. Other aspects of its reproduction, as well as its tadpoles, remain undescribed as of 2008. Though tiny and inconspicuous, it is not considered a particularly rare species. With a considerable range including several protected areas and apparently able to live in secondary forest and tolerate some amount of human use of its habitat, it is not considered threatened by the IUCN. Systematics and taxonomy This frog has a complicated taxonomic history: it was initially described in the genus Centrolenella, which is nowadays included in Centrolene. In 2009, it was transferred further to Vitreorana. In 2008, it was found to include that the frogs that had been described as Centrolenella ametarsia are indistinguishable from the present species. The holotype of this species is specimen MNHNP 1973.1673, that of C. ametarsia is specimen MCZ A96522.