relationships, thus Incilius should be treated as a subgenus of Bufo. Frost refuted these criticisms in a 2009 response. A phylogenetic analysis by Mendelson based on morphology, life history, and molecular data was published 2011 which recommending sinking Cranopsis/Cranophryne, Ollotis and Crepidius/Crepidophryne back into Incilius. Mendelson et al. (2011) suggest that this species is part of an I. valliceps species group, or more specifically in what they termed the "Forest Group" subgroup, including the taxa I. aucoinae, I. cavifrons, I. campbelli, I. cristatus, I. leucomoyos, I. macrocristatus, I. spiculatus and I. tutelarius. Description Incilius melanochlorus are moderately large toads, with males attaining to in snout–vent length, and females to . There is marked sexual dimorphism, the females having more irregular dark colouration on their back and being larger, but the males having longer heads and more brawny forearms. The head is wider than it is long. The head is crested, with the cranium sporting well-developed, thin and high crests. This crest ornamentation consists of canthal, supraorbital, and postpreorbital crests, and parietal crests with transverse folds between them (other species have crests on other locations). The crests run from the nostril, behind the eye, to the back of the head. The skin of the top of the head is hardened and stuck to the top of the skull. Males have a vocal sac. The vocal slits which hide the vocal sac inside the throat are small and bilateral, the sac has a single lobe, and is large, heavily pigmented with a greenish cast, and when calling inflates fully to a round shape. The eyes are large with a coppery bronze iris and a black pupil, the top half of the iris lighter than the bottom half, with a thin, dark line separating the two halves. The tongue is long and thin. The paratoid glands are small, smaller than the upper eyelid, and triangular in shape, and the toad has a small tympanum. The hind legs are relatively short, as are the feet and the tibia, but the toes are long and have some webbing between them. The first finger is longer than the second. The fingers and toes have tubercles underneath them. Sometimes the tips of the fingers and toes are lighter-coloured, sometimes the same colour as the rest of the digit. The tarsus (heel) has two differently-shaped tubercles, one facing inwards, the other outwards. The dorsum (back) is very warty,