calls and between the pulses also decrease, whereas the pulse rate increases. The number of pulses per call and the duration of the pulses do not change. Annual and daily calling activity Calling of the males begins in the first half of February and lasts until October. It reaches a climax in spring and early summer. During this time calling begins shortly after sunset and lasts until 1 to 2 a.m. Later in summer calling ends earlier, around 10 to 11 p.m. and ends in September and October even earlier. Calling behaviour Calling males gather in small shallow ponds, irrigation canals and ditches, rice and corn fields. They form choruses consisting of about 20 males. The calling males maintain a minimum distance of about 0.7 meters. At decreasing calling activity during the year, males continue to form choruses, but the number of choruses decreases. Males call while sitting in the water, the body is erected, the hind limbs and the posterior part of the body are immersed in the water. The males have a single subgular vocal sac. When fully inflated, it is very large in relation to the body length. Pair formation is high in spring and early summer, later on it decreases, but pairs are observed until October. Status Very little is known about this species and its natural history. It inhabits marshy areas and presumably its larvae are aquatic. Its population status, ecology and the threats it faces are unknown and the International Union for Conservation of Nature lists its conservation status as "data deficient". References vittata Frogs of Africa Amphibians of Uganda Endemic fauna of Uganda Taxa named by George Albert Boulenger Amphibians described in 1906 Taxonomy articles created by Polbot