Tilda Johnson, introduced as the Queen of the Werewolves and also known as Dr. Nightshade, Deadly Nightshade, or simply Nightshade, is a fictional character appearing in American comic books published by Marvel Comics. Introduced as a supervillain opposing Captain America, Falcon, Power Man, Iron Fist, and Black Panther, she is later reformed, becoming the superhero Nighthawk and joining the Avengers in 2017. Gabrielle Dennis portrayed Tilda Johnson in the second season of the Marvel Cinematic Universe (MCU) / Netflix series Luke Cage. Additionally, Nabiyah Be was intended to portray Johnson in the MCU film Black Panther, but was renamed Linda Johnson due to Dennis being announced to portray Johnson in Luke Cage. Publication history Tilda Johnson first appeared in Captain America and the Falcon #164 (August 1973), created by Steve Englehart and Alan Lee Weiss. Fictional character biography Queen of the Werewolves Tilda Johnson was born into poverty in New York City. At an early age, she discovered that she had a natural aptitude for science. As a teenager, she used her extensive knowledge to begin a career as a criminal scientist. The Yellow Claw helped her develop a method for turning normal humans into obedient werewolf-like creatures, and attempted to transform convicts into a werewolf army, but the pair were defeated by Captain America and S.H.I.E.L.D. She later took control of a S.H.I.E.L.D. facility, and used pheromones to force Captain America to battle the Falcon, but was defeated. She clashes with Power Man and Iron Fist on several occasions. Nightshade later appeared as second-in-command of Superia's Femizons. She helps develop a serum to transform men into women which she uses on Captain America and Paladin. She also helped Superia develop her sterilization bomb. She was also involved in a scheme with Dredmund Druid, though as a double agent for Superia. She used a serum to transform Captain America and the ordinary people of Starkesboro, Massachusetts into pseudo-werewolves. MODOK's 11 Nightshade joined MODOK's 11, with the main objective of stealing the powerful Hypernova. It has been revealed after her last battle against the Black Panther, she tried to start her life over without crime. However, as she was self-taught and had no official degrees, the only medical job she could get was as a receptionist at a hospital. After she pointed out a mistake by one of the doctors, she was fired immediately and contemplated suicide, until MODOK contacted her