an impressive amount in their children before they are even born. The time and nutrients required to develop the fetus, and the risks associated with both giving these nutrients and undergoing childbirth, are a sizable investment. To ensure that this investment is not for nothing, mothers are likely to invest in their children after they are born, to be sure that they survive and are successful. Relative to most other species, human mothers give more resources to their offspring at a higher risk to their own health, even before the child is born. This is associated with the evolution of a slower life history, in which fewer, larger offspring are born after longer intervals, requiring increased parental investment. The developing human fetus––and especially the brain––requires nutrients to grow. In the later weeks of gestation, the fetus requires increasing nutrients as the growth of the brain increases. Rodents and primates have the most invasive placenta phenotype, the hemochorial placenta, in which the chorion erodes the uterine epithelium and has direct contact with maternal blood. The other placental phenotypes are separated from the maternal bloodstream by at least one layer of tissue. The more invasive placenta allows for a more efficient transfer of nutrients between the mother and fetus, but it comes with risks as well. The fetus is able to release hormones directly into the mother's bloodstream to “demand” increased resources. This can result in health problems for the mother, such as pre-eclampsia. During childbirth, the detachment of the placental chorion can cause excessive bleeding. The obstetrical dilemma also makes birth more difficult and results in increased maternal investment. Humans have evolved both bipedalism and large brain size. The evolution of bipedalism altered the shape of the pelvis, and shrunk the birth canal at the same time brains were evolving to be larger. The decreasing birth canal size meant that babies are born earlier in development, when they have smaller brains. Humans give birth to babies with brains 25% developed, while other primates give birth to offspring with brains 45-50% developed. A second possible explanation for the early birth in humans is the energy required to grow and sustain a larger brain. Supporting a larger brain gestationally requires energy the mother may be unable to invest. The obstetrical dilemma makes birth challenging, and a distinguishing trait of humans is the need for assistance during childbirth. The altered shape of the bipedal