in the protected area is the Northwest African cheetah (Acinonyx jubatus hecki). As of 2007, there were only about 5–13 individuals left in the national park and neighboring W National Park. By 2009, the lion (Panthera leo leo) population in the WAP Complex comprised about 100 animals and was possibly the largest in West Africa at the time. The endangered West African wild dog (Lycaon pictus manguensis) was recorded in Pendjari National Park during a survey in April 2000, as well as African leopard, spotted hyena, side-striped jackal, and African civet. The number of waterbucks (Kobus ellipsiprymnus defassa) decreased from about 3,000 in the 1970s to 120 in 2004. Birds The hooded vulture (Necrosyrtes monachus) is resident in the protected area, but in small numbers. The park is renowned for its abundance of bird species. Some 300 different species are present. Pallid harrier (Circus macrourus) and lesser kestrel (Falco naumanni) are occasionally recorded and there are a few isolated records for lappet-faced vulture (Torgos tracheliotus). Fox kestrel (Falco alopex) is not uncommon, while the African swallow-tailed kite (Chelictinia riocourii) is a not uncommon dry season visitor. The booted eagle (Hieraaetus pennatus) has also been recorded here. BirdLife notes that "the Pendjari is notable for large conspicuous species such as African openbill stork (Anastomus lamelligerus), Abdim's stork (Ciconia abdimii), saddle-billed stork (Ephippiorhynchus senegalensis), and seasonally, flocks of up to 60 European white storks (Ciconia ciconia). The African fish eagle (Haliaeetus vocifer) and Pel's fishing-owl (Scotopelia peli) can also be found." Among the more notable species recorded are pied-winged swallow (Hirundo leucosoma), white-crowned robin-chat (Cossypha albicapillus), Botta's wheatear (Oenanthe bottae), familiar chat (Cercomela familiaris), white-fronted black-chat (Myrmecocichla albifrons), mocking cliff-chat (Thamnolaea cinnamomeiventris), common rock thrush (Monticola saxitilis), Senegal eremomela (Eremomela pusilla), blackcap babbler (Turdoides reinwardtii), red-winged pytilia (Pytilia phoenicoptera), black-rumped waxbill (Estrilda troglodytes), bush petronia (Petronia dentata) and Togo paradise-whydah (Vidua togoensis). Grey tit-flycatcher (Myioparus plumbeus) has been recorded as well as several other species of the undergrowth. White-throated greenbul (Phyllastrephus albigularis) has been recorded at Tanguiéta and the white-throated francolin (Francolinus albogularis), a rare resident, has been spotted in farmland south of Natitingou. South of the park there is a large semi-protected zone known in French as La zone cygnetique de la Pendjari where a number of other species have been spotted. The National park and the bird habitat is protected by the government in Benin. Other bird species include: Fox