until 1900. 1865 to 1900 saw a time of theory formulation in the field of heredity/genetics. In 1883, Wilhelm Roux argued that the linear structure of chromosomes has an impact of making sure daughter cells get equal amounts of chromosomal material. This was the beginning of the chromosome theory; Roux viewed his findings as argument that chromosomes contain units of heredity. During this time frame, Hugo de Vries put forth a theory that persistent hereditary units are passed through generations and that each “unit” deals with a specific characteristic and the units can combine in different ways in the offspring. From 1900 to 1909, anomalous data began to accumulate. Gene linkage was first reported by Carl Correns in 1900, contradicting Mendel's law of independent assortment. Thomas Hunt Morgan was the first to provide a working hypothesis for these exceptions. He postulated that genes that remained together while being passed from generation to generation must be located on the same chromosome. Sturtevant's work and its importance Sturtevant's most notable discoveries include the principle of genetic mapping, the first reparable gene defect, the principle underlying fate mapping, the phenomena of unequal crossing-over, and position effect. His main contributions to science include his analysis of genetic “linkage groups,” which became a classical method of chromosome mapping that we still use today. In 1913, he determined that genes were arranged on chromosomes in a linear fashion, like beads on a necklace. He also showed that the gene for any specific trait was in a fixed location (locus). In his work between 1915 and 1928, Sturtevant determined that genes of Drosophila are arranged in linear order. In 1920, he published a set of three papers under the title “Genetic Studies on Drosophila simulans,” which “proved that two closely related species had newly recurring mutations that were allelic and thus probably identical” (Provine 2). His work also helped to determine the chromosomal basis of sex determination and development and described the importance of chromosomal crossing-over or recombination in genetic linkage of traits. One of Sturtevant's principal contributions was his introduction of the concept that the frequency of crossing-over between two genes could help determine their proximity on a linear genetic map. His experiments determined that the frequency of double crossing over can be used to deduce gene order. He demonstrated this concept by constructing crosses of three segregating genes, called "three-factor crosses". He found that using