forester and political activist, and Emily Morgan, a hairdresser and shop owner. With his partner, Jette, he shares five children (plus spouses) and eight grandchildren, of whom five are adopted. Experimental contributions In bacteria: With M. Meselson, the demonstration of semiconservative DNA replication. In phage T4: With H. Foss and others, demonstrations of genetic linkage circularity and its relation to genetic heterozygosis. With N. Murray and others, the determination, by genetic methods, of the direction of mRNA synthesis on cotranscribed pairs of genes. In Lambda: With M. Stahl and others, the discovery and analysis of the genetic element, Chi, that stimulates nearby genetic recombination in bacteria. With M. Stahl and others, the mutual dependence of DNA replication and genetic recombination. These studies utilized the method of density gradient centrifugation that was developed for the test of the semiconservative model of DNA replication. In Yeast: With H. Foss and others, the demonstration of two functional pathways for genetic recombination in wild-type budding yeast. Theoretical contributions With C. Steinberg, formulations of phage growth, recombination and mutation. With J. Szostak and others, the interpretation of genetic recombination in terms of the repair of double-strand DNA breaks. With R. Lande, E. Housworth and others, mathematical formalizations of recombination in higher organisms. Selected honors 1997- Fellow, American Academy of Microbiology 1996 Thomas Hunt Morgan Medal (from Genetics Society of America) 1986- Associate Member EMBO 1985- American Cancer Society Research Professor 1985-1990 MacArthur Fellow 1981- Member, American Academy of Arts and Sciences 1976- Member, National Academy of Sciences 1975-76; 1985-1986 Guggenheim Fellow 1969-70 NIH Special Postdoctoral Fellowship Honorary Doctor of Science: Oakland University and University of Rochester References External links Franklin Stahl in the Encyclopædia Britannica Franklin Stahl at DNAi.org Franklin Stahl at the University of Oregon Animation of the Meselson-Stahl Experiment from the Dolan DNA Learning Center Description of the Meselson-Stahl Experiment written by Nathan H. Lents, including original data from Visionlearning 1929 births Members of the United States National Academy of Sciences Living people American molecular biologists History of genetics MacArthur Fellows Phage workers University of Oregon faculty University of Missouri faculty American biochemists Harvard College alumni University of Rochester alumni