January 2013 and March 2018. In Bhutan, it has been recorded in Royal Manas National Park, and in broadleaved and mixed conifer forests at elevations up to in Jigme Dorji National Park and Wangchuck Centennial National Park. In Thailand, it was recorded in a hill evergreen bamboo mixed forest in Phu Khieu Wildlife Sanctuary. In Borneo, it has also been recorded in peat swamp forest. The population size of the marbled cat is not well understood. Few records were obtained during camera-trapping surveys throughout much of its range. In three areas in Sabah, the population density was estimated at 7.1 to 19.6 individuals per , an estimate that may be higher than elsewhere in the cat's range. In Kalimantan, marbled cats were recorded in mixed swamp forest and tall interior forest at altitudes below in the vicinity of Sabangau National Park between 2008 and 2018. Behaviour and ecology Marbled cats recorded in northeastern India and Kalimantan on Borneo were active by day. The first-ever radio-tracked marbled cat had an overall home range of at an elevation of and was active primarily during nocturnal and crepuscular times. Marbled cats recorded in northeast India were active during the day with activity peaks around noon. Forest canopies probably provide the marbled cat with much of its prey: birds, squirrels and other rodents, and reptiles. In the Bukit Barisan Selatan National Park, a marbled cat was observed in a dense forest patch in an area also used by siamang. In Thailand, one individual has been observed in Phu Khieo Wildlife Sanctuary preying on a Phayre's leaf monkey. A few marbled cats have been bred in captivity, with gestation estimated to be 66 to 82 days. In the few recorded instances, two kittens were born in each litter, and weighed from . Their eyes open at around 12 days, and the kittens begin to take solid food at two months, around the time that they begin actively climbing. Marbled cats reach sexual maturity at 21 or 22 months of age, and have lived for up to 12 years in captivity. Threats Indiscriminate snaring is prevalent throughout much of its range, and likely poses a major threat. It is valued for its skin, meat, and bones, but infrequently observed in the illegal Asian wildlife trade. During a survey in the Lower Subansiri District of Arunachal Pradesh, a marbled cat was encountered that had been killed by