plots like many of its predecessors (and the various contenders within the family sought power and often succeeded in overthrowing the preceding kinsman); Alexios II, the first Komnenos to ascend as a minor, ruled for three years and his conqueror and successor Andronikos I ruled for two, overthrown by the Angelos family under Isaac II who was dethroned and blinded by his own brother Alexios III. The Angeloi were overthrown during the Fourth Crusade in 1204, by Alexios V Doukas, a relative from the Doukas family. Later family Several weeks before the occupation of Constantinople by crusaders in 1204, one branch of the Komnenoi fled back to their homelands in Paphlagonia, along the eastern Black Sea and its hinterland in the Pontic Alps, where they established the Empire of Trebizond. Their first 'emperor', named Alexios I, was the grandson of Emperor Andronikos I. These emperors – the Grand Komnenoi ( or in Greek) as they were known – ruled in Trebizond for over 250 years, until 1461, when David Komnenos was defeated and executed by the Ottoman sultan Mehmed II. Mehmed himself claimed descent from the Komnenos family via John Tzelepes Komnenos. The Trapezutine branch of the Komnenos dynasty also held the name of Axouchos as descendants of John Axouch, a Byzantine nobleman and minister to the Byzantine Komnenian Dynasty. A princess of the Trebizond branch is said to have been the mother of prince Yahya (born 1585), who reportedly became a Christian yet spent much of his life attempting to gain the Ottoman throne. Another branch of the family, descendants of Constantine Angelos, founded the Despotate of Epirus in 1204, under Michael I Komnenos Doukas, great-grandson of Emperor Alexios I. This branch adopted the surnames Komnenos Doukas and are known as such in modern scholarship. Helena Doukaina Komnene, a child of that branch of the family, married Guy I de la Roche thereby uniting the Komnenos and the de la Roche houses, with Komnenos family members eventually becoming Dukes of Athens. One renegade member of the family, also named Isaac Komnenos, established a separate "empire" on Cyprus in 1184, which lasted until 1191, when the island was taken from him by Richard I of England during the Third Crusade. His daughter, called the Damsel of Cyprus, married Thierry of Flanders during the Fourth Crusade and tried to claim the island. When the Byzantine Empire was restored in 1261 at