wider. They were found as far north as the Mediterranean coast in the Nile Delta and across the Red Sea in Israel, Palestine and Syria. The Nile crocodile has historically been recorded in areas where they are now regionally extinct. For example, Herodotus recorded the species inhabiting Lake Moeris in Egypt. They are thought to have become extinct in the Seychelles in the early 19th century (1810–1820). Today, Nile crocodiles are widely found in, among others, Somalia, Ethiopia, Uganda, Kenya, Egypt, the Central African Republic, the Democratic Republic of the Congo, Equatorial Guinea, Tanzania, Rwanda, Burundi, Zambia, Zimbabwe, Gabon, Angola, South Africa, Malawi, Mozambique, Namibia, Sudan, South Sudan, Botswana, and Cameroon. The Nile crocodile's current range of distribution extends from the regional tributaries of the Nile in Sudan and Lake Nasser in Egypt to the Cunene river of Angola, the Okavango Delta of Botswana, and the Olifants river in South Africa. Isolated populations also exist in Madagascar, which likely colonized the island after the extinction of the endemic crocodile Voay. In Madagascar, crocodiles occur in the western and southern parts from Sambirano to Tôlanaro. They have been spotted in Zanzibar and the Comoros in modern times, but occur very rarely. The species was previously thought to extend in range into the whole of West and Central Africa, but these populations are now typically recognized as a distinct species, the West African (or desert) crocodile. The distributional boundaries between these species were poorly understood, but following several studies, they are now better known. West African crocodiles are found throughout much of West and Central Africa, ranging east to South Sudan and Uganda where the species may come into contact with the Nile crocodile. Nile crocodiles are absent from most of West and Central Africa, but range into the latter region in eastern and southern Democratic Republic of Congo, and along the Central African coastal Atlantic region (as far north to Cameroon). Likely a level of habitat segregation occurs between the two species, but this remains to be confirmed. Nile crocodiles may be able to tolerate an extremely broad range of habitat types, including small brackish streams, fast-flowing rivers, swamps, dams, and tidal lakes and estuaries. In East Africa, they are found mostly in rivers, lakes, marshes, and dams, favoring open, broad bodies of water over smaller ones. They are often found in waters adjacent to various open habitats such as savanna