local collectors in the Chapada do Araripe, especially on fish fossils. The buyers were unaware of the modifications to the specimen until it was sent to universities in the United Kingdom for CT scan imaging. This revealed the collectors had tried to reconstruct the skull by grafting parts of the maxilla (main upper jaw bone) onto the front of the rostrum (snout). The skull (designated SMNS 58022) became the holotype specimen of the new genus and species Irritator challengeri in February 1996, when it was first scientifically described by paleontologists David M. Martill, Arthur R.I. Cruickshank, Eberhard Frey, Philip G. Small and Maria Clarke. In this paper, Martill and his team wrote that the generic name Irritator came "from irritation, the feeling the authors felt (understated here) when discovering that the snout had been artificially elongated." The type species, Irritator challengeri, was named after Professor Challenger, a character in Arthur Conan Doyle's novels, specifically The Lost World. Two years earlier, Frey and Martill had named a new pterosaur species from the Crato Formation Arthurdactylus conandoylei, after the novelist himself. When Martill and colleagues first described Irritator challengeri, the holotype was still extensively encased in calcareous rock matrix. Under the supervision of American paleontologist Hans-Dieter Sues, technician Diane M. Scott from the University of Toronto at Mississauga assumed the task of fully extracting the skull bones from the rock, allowing for a detailed redescription in 2002. Published by Sues, Frey, Martill, and Scott, this inspection of the now fully prepared specimen negated many of Martill and colleagues' original observations, which were based on misinterpretations of the damaged and largely concealed skull. The estimated length of the complete skull was shorter than previously proposed. What was originally thought to be a prominent head crest proved to be an unattached, indeterminate bone fragment. As in the previous study, Sues and colleagues regarded the African genus Spinosaurus as the most similar taxon to Irritator, because they shared many dental features, including mostly straight conical tooth crowns, thin enamel, well-defined edges with no , and lengthwise fluting. Since little was known of Spinosauruss skull at the time, these similarities were enough for the authors to suggest a possible junior synonymy of Irritator with Spinosaurus. Sues and colleagues noted that more overlapping skull material was needed for further diagnosis. As more of Spinosauruss skull became known, later research maintained separation of the two taxa. Although the