Some objection has been raised to these assertions. Kellner and Campos in 2000 and Brazilian paleontologist Elaine B. Machado and Kellner in 2005 expressed the opinion that the fossils come from two different genera, and that the holotype of Angaturama limai was clearly more laterally flattened than that of Irritator challengeri. A review of both fossils by the Brazilian paleontologists Marcos A. F. Sales and Cesar L. Schultz in 2017 noted that the specimens also differ in other aspects of their preservation: the Irritator specimen is brighter in color and is affected by a vertical crack, while the Angaturama specimen bears many cavities; the damage to the teeth of the Irritator challengeri holotype is also much less severe. Sales and Schultz also identified a possible point of overlap, the third left maxillary tooth, and observed that the skull of Angaturama could have been larger than that of Irritator based on the proportions of the closely related genus Baryonyx. They therefore concluded that the two specimens do not belong to the same individual, Sales and Schultz noted that synonymy at the genus level would need to be verified by more extensively overlapping remains. If Angaturama and Irritator are regarded as a member of the same genus, the latter would be the valid scientific name under rules of priority, since it was named almost a month earlier. Postcranial material and additional finds Besides the skull, the snout fragment, and some isolated teeth, the Romualdo Formation has also yielded l remains that may belong to spinosaurids, many of which are hitherto undescribed, and all of them pertaining to the Spinosaurinae subfamily. In 2004, parts of a spinal column (MN 4743-V) were unearthed at the formation. Brazilian paleontologist Jonathas Bittencourt and Kellner assigned these, due to their structure, to the Spinosauridae. It is uncertain whether this specimen can be referred to Irritator or Angaturama, given that both are based only on skull material. In 2007, Machado and Kellner tentatively referred a rib fragment (MN 7021-V) to the Spinosauridae. However, the most complete spinosaur specimen retrieved from the Romualdo Formation is MN 4819-V, a partial skeleton lacking the skull. First reported in 1991, the specimen was referred by Kellner to the Spinosauridae in 2001 because of its tall sacral neural spines and the enlarged condition of the hand claw. The skeleton was fully described in 2010 in an as-of-yet unpublished master's thesis by Machado. An