When Reginald Innes Pocock reviewed Felis skins in the collection of the Natural History Museum, London, he considered Felis reyi a synonym of Felis lybica sarda, the Sardinian wild cat. The Corsican wildcat is considered to have been introduced in the early first millennium. The earliest known fossil records of cats date to the early 14th century, but older chronostratigraphic layers revealed fossils of livestock introduced since the Iron Age. Felis lybica jordansi proposed by Ernst Schwarz in 1930 was a skull and skin of a male specimen from Santa Margarita in Mallorca that had more pronounced stripes than the African wildcat. This is also considered to have descended from domestic cats introduced to the island. Felis silvestris cretensis proposed by Theodor Haltenorth in 1953 was a cat skin purchased in a bazaar in Chania that resembled an African wildcat, but had a bushy tail like a European wildcat. Groves considered the Cretan wildcat an introduced feral cat. Distribution and habitat The feral cat is the most widely distributed terrestrial carnivore. It occurs between 55° North and 54.3° South latitudes in a wide range of climatic zones and islands in the Atlantic, Indian and Pacific Oceans, and the Mediterranean Sea, including Canary Islands, Port-Cros, Dassen Island, Marion Island, Juan de Nova Island, Réunion, Hahajima, Okinawa Island, Raoul Island, Herekopare Island, Stewart Island, Macquarie Island, Galápagos Islands, San Clemente Island, Isla Natividad, San José Island, and New Island. Feral cat colonies also occur on the Japanese islands of Ainoshima, Hahajima and Aoshima, Ehime. The feral cat population on the Hawaiian Islands is mainly of European origin and probably arrived in the 19th century on ships. Feral cat colonies in Rome have been monitored since 1991. Urban feral cats were studied in Madrid, Jerusalem and Ottawa. Behavior and ecology Some behaviors of feral cats are commonly observed, although there is disagreement among veterinarians, rescuers and researchers on the prevalence of some. In a free-roaming environment, feral cats avoid humans. They do not allow themselves to be handled or touched by humans, and back away or run when they are able to do so. If trapped, they hiss, growl, bare their teeth, or strike out. They remain fairly hidden from humans and will not approach, although some feral cats gradually become more comfortable around humans who feed them regularly. Most feral cats have small home ranges, although some are more transient and travel