largest in the world in terms of the number of languages (1,436) it contains. The vast majority of languages of this family are tonal such as Yoruba, and Igbo, However, others such as Fulani, Wolof and Kiswahili are not. A major branch of the Niger–Congo languages is Bantu, which covers a greater geographic area than the rest of the family. Bantu speakers represent the majority of inhabitants in southern, central and southeastern Africa, though San, Pygmy, and Nilotic groups, respectively, can also be found in those regions. Bantu-speakers can also be found in parts of Central Africa such as the Gabon, Equatorial Guinea and southern Cameroon. Swahili, a Bantu language with many Arabic, Persian and other Middle Eastern and South Asian loan words, developed as a lingua franca for trade between the different peoples in southeastern Africa. In the Kalahari Desert of Southern Africa, the distinct people known as Bushmen (also "San", closely related to, but distinct from "Hottentots") have long been present. The San evince unique physical traits, and are the indigenous people of southern Africa. Pygmies are the pre-Bantu indigenous peoples of Central Africa. Nilo-Saharan The Nilo-Saharan languages are concentrated in the upper parts of the Chari and Nile rivers of Central Africa and Southeast Africa. They are principally spoken by Nilotic peoples and are also spoken in Sudan among the Fur, Masalit, Nubian and Zaghawa peoples and in West and Central Africa among the Songhai, Zarma and Kanuri. The Old Nubian language is also a member of this family. Major languages of Africa by region, family and number of primary language speakers in millions: Genetic history Major cities Sub-Saharan Africa has several large cities. Lagos is a city in the Nigerian state of Lagos. The city, with its adjoining conurbation, is the most populous in Nigeria, and the second-most populous in Africa after Cairo, Egypt. It is one of the fastest-growing cities in the world, and also one of the most populous urban agglomerations. Lagos is a major financial centre in Africa; this megacity has the highest GDP, and also houses Apapa, one of the largest and busiest ports on the continent. Dar es Salaam is the former capital of, as well as the most populous city in, Tanzania; it is a regionally important economic centre. It is located on the Swahili coast. Johannesburg is the largest city in South Africa. It is the provincial capital and