only the subfamily Trichechinae (Potamosiren, Ribodon, and Trichechus), but also Miosireninae (Anomotherium and Miosiren). The African manatee and the West Indian manatee are more closely related to each other than to the Amazonian manatee. Dugongidae comprises the subfamilies Dugonginae and Hydrodamalinae and the paraphyletic Halitheriinae. The tusks of modern-day dugongs may have originally been used for digging, but they are now used for social interaction. The genus Dugong probably originated in the Indo-Pacific. Description Adaptations The tail fluke of a dugong is notched and similar to those of dolphins, whereas the tail fluke of manatee is paddle-shaped. The fluke is pumped up and down in long strokes to move the animal forward, or twisted to turn. The forelimbs are paddle-like flippers which aid in turning and slowing. Unlike manatees, the dugong lacks nails on its flippers, which are only 15% of a dugong's body length. Manatees generally glide at speeds of , but can reach speeds of in short bursts. The body is fusiform to reduce drag in the water. Like those of cetaceans, the hind limbs are internal and vestigial. The snout is angled downwards to aid in bottom-feeding. Sirenians typically make two- to three-minute dives, but manatees can hold their breath for up to 15 minutes while resting and dugongs up to six minutes. They may stand on their tails to hold their heads above water. Much like elephants, manatees are polyphyodonts, continuously replacing their teeth from the back of the jaw. Adults lack incisors, canines, and premolars, and instead have 8 to 10 cheek teeth. Manatees have an unlimited supply of teeth moving in from the back and shedding in the front; these are continuously formed by a dental capsule behind the tooth row. These teeth are constantly worn down by the abrasive vascular plants they forage, particularly aquatic grasses. Unlike those of manatees, the dugong's teeth do not continually grow back via horizontal tooth replacement. The dugong has two tusks which emerge in males during puberty, and sometime later in life for females after reaching the base of the premaxilla. The number of growth layer groups in a tusk indicates the age of a dugong. Sirenians exhibit pachyostosis, a condition in which the ribs and other long bones are solid and contain little or no bone marrow. They have among the densest bones in the animal kingdom. These may act as ballast, countering the buoyancy of