of the northern Ozarks, the Wabash basin, the Maumee River drainage, and along the shores of Lake Erie and Lake Ontario. Etymology Etheostoma: meaning filter or strain and meaning mouth E. b. blennioides: referring to its resemblance to the Mediterranean blennies Constantine S. Rafinesque knew in his early years E. b. newmanii: patronymic for Francis H. Newman, aquatic biologist, who collected the type specimen E. b. pholidotum: meaning scaled, referring to its fully scaled belly. Habitat Greenside darters inhabit gravel riffles of large creeks to medium rivers and often are found in swift waters over large boulders and large rubble. Systematics The greenside darter was first formally described in 1819 by the French naturalist Constantine Samuel Rafinesque (1783-1840) with the type locality given as the Ohio River. Rafinesque placed the new species in a new genus Etheostoma and it was subsequently designated as the type species of that genus by Louis Agassiz in 1854. Rafinesque gave the species the specific name blennoidies because of its resemblance to the blennies. As previously stated, Miller (1968) concluded the E. blennioides complex consists of the four subspecies above. Unpublished studies by Richard L. Mayden and colleagues indicate E. b. newmanii from the Cumberland and Tennessee Rivers may be a valid species and distinct from other members of the complex. The former subspecies, E. b. gustelli has recently been re-elevated to species status, Etheostoma gustelli (Tuckasegee darter) based on lack of hybridization. Etheostoma blennioides belongs to the subgenus Etheostoma that contains these species: E. blennius (blenny darter) E. gutselli (Tuckasegee darter) E. histrio (harlequin darter) E. inscriptum (turquoise darter) E. lynceum (brighteye darter) E. rupestre (rock darter) E. sellare (Maryland darter) E. swannanoa (Swannanoa darter) E. thalassinum (seagreen darter) E. zonale (banded darter) Etheostoma blennioides is further nested within the greenside darter group that contains E. blennius, E. gutselli, E. inscriptum, E. swannanoa, and E. thalassinum. References Etheostoma Fish described in 1819 Taxa named by Constantine Samuel Rafinesque Freshwater fish of North America