Permoplectoptera still had 9 pairs of abdominal gills, and the adults still had long hindwings. Maybe the fossil family Cretereismatidae from the Lower Cretaceous Crato Formation of Brazil also belongs as the last offshoot to Permoplectoptera. The Crato outcrops otherwise yielded fossil specimens of modern mayfly families or the extinct (but modern) family Hexagenitidae. However, from the same locality the strange larvae and adults of the extinct family Mickoleitiidae (order Coxoplectoptera) have been described, which represents the fossil sister group of modern mayflies, even though they had very peculiar adaptations such as raptorial forelegs. The oldest mayfly inclusion in amber is Cretoneta zherichini (Leptophlebiidae) from the Lower Cretaceous of Siberia. In the much younger Baltic amber numerous inclusions of several modern families of mayflies have been found (Ephemeridae, Potamanthidae, Leptophlebiidae, Ametropodidae, Siphlonuridae, Isonychiidae, Heptageniidae, and Ephemerellidae). The modern genus Neoephemera is represented in the fossil record by the Ypresian species N. antiqua from Washington state. Grimaldi and Engel, reviewing the phylogeny in 2005, commented that many cladistic studies had been made with no stability in Ephemeroptera suborders and infraorders; the traditional division into Schistonota and Pannota was wrong because Pannota is derived from the Schistonota. The phylogeny of the Ephemeroptera was first studied using molecular analysis by Ogden and Whiting in 2005. They recovered the Baetidae as sister to the other clades. Mayfly phylogeny was further studied using morphological and molecular analyses by Ogden and others in 2009. They found that the Asian genus Siphluriscus was sister to all other mayflies. Some existing lineages such as Ephemeroidea, and families such as Ameletopsidae, were found not to be monophyletic, through convergence among nymphal features. The following traditional classification is based on Peters and Campbell (1991), in Insects of Australia. Suborder Pannota Superfamily Ephemerelloidea Ephemerellidae Leptohyphidae Tricorythidae Superfamily Caenoidea Neoephemeridae Baetiscidae Caenidae Prosopistomatidae Suborder Schistonota Superfamily Baetoidea Siphlonuridae Baetidae Oniscigastridae Ameletopsidae Ametropodidae Superfamily Heptagenioidea Coloburiscidae Oligoneuriidae Isonychiidae Heptageniidae Superfamily Leptophlebioidea Leptophlebiidae Superfamily Ephemeroidea Behningiidae Potamanthidae Euthyplociidae Polymitarcyidae Ephemeridae Palingeniidae Phylogeny After In human culture In art The Dutch Golden Age author Augerius Clutius (Outgert Cluyt) illustrated some mayflies in his 1634 De Hemerobio ("On the Mayfly"), the earliest book written on the group. Maerten de Vos similarly illustrated a mayfly in his 1587 depiction of the fifth day of creation, amongst an assortment of fish and water birds. In 1495 Albrecht Dürer included a mayfly in his engraving The Holy Family with