Landscape ecology is the science of studying and improving relationships between ecological processes in the environment and particular ecosystems. This is done within a variety of landscape scales, development spatial patterns, and organizational levels of research and policy. Concisely, landscape ecology can be described as the science of "landscape diversity" as the synergetic result of biodiversity and geodiversity. As a highly interdisciplinary field in systems science, landscape ecology integrates biophysical and analytical approaches with humanistic and holistic perspectives across the natural sciences and social sciences. Landscapes are spatially heterogeneous geographic areas characterized by diverse interacting patches or ecosystems, ranging from relatively natural terrestrial and aquatic systems such as forests, grasslands, and lakes to human-dominated environments including agricultural and urban settings. The most salient characteristics of landscape ecology are its emphasis on the relationship among pattern, process and scale, and its focus on broad-scale ecological and environmental issues. These necessitate the coupling between biophysical and socioeconomic sciences. Key research topics in landscape ecology include ecological flows in landscape mosaics, land use and land cover change, scaling, relating landscape pattern analysis with ecological processes, and landscape conservation and sustainability. Landscape ecology also studies the role of human impacts on landscape diversity in the development and spreading of new human pathogens that could trigger epidemics. Terminology The German term Landschaftsökologie–thus landscape ecology–was coined by German geographer Carl Troll in 1939. He developed this terminology and many early concepts of landscape ecology as part of his early work, which consisted of applying aerial photograph interpretation to studies of interactions between environment and vegetation. Explanation Heterogeneity is the measure of how parts of a landscape differ from one another. Landscape ecology looks at how this spatial structure affects organism abundance at the landscape level, as well as the behavior and functioning of the landscape as a whole. This includes studying the influence of pattern, or the internal order of a landscape, on process, or the continuous operation of functions of organisms. Landscape ecology also includes geomorphology as applied to the design and architecture of landscapes. Geomorphology is the study of how geological formations are responsible for the structure of a landscape. History Evolution of theory One central landscape ecology theory originated from MacArthur & Wilson's The Theory of Island Biogeography. This work considered the biodiversity on islands as the result of competing forces of colonization from a mainland stock and stochastic extinction. The concepts of