Aphodius is a genus of beetles in the family Scarabaeidae. In most species both the adults and larvae are coprophagous (dung feeding) although some species have herbivorous or saprophagous larvae. Aphodius species typically dominate dung beetle communities in north temperate ecosystems. Most species are functionally classified as endocoprids, also known as dwellers, because the larvae live and feed within the dung pat itself. Species These 44 species belong to the genus Aphodius, including 21 extinct species. Aphodius beloni Mulsant & Godart, 1879 Aphodius calichromus Balthasar, 1932 Aphodius cardinalis Reitter, 1892 Aphodius clypeatus Fischer von Waldheim, 1821 Aphodius coniugatus (Panzer, 1795) Aphodius corallifer Koshantschikov, 1913 Aphodius crux Wiedemann, 1823 Aphodius elegans Allibert, 1847 Aphodius fasciger Harold, 1881 Aphodius fimetarius (Linnaeus, 1758) Aphodius foetidus (Herbst, 1783) Aphodius guangdongensis Maté, 2008 Aphodius irregularis Westwood, 1839 Aphodius marginatus Fischer von Waldheim, 1842 Aphodius micros Walker, 1871 Aphodius minatorius Péringuey, 1908 Aphodius pedellus (Degeer, 1774) Aphodius pedrosi Wollaston, 1854 Aphodius plasoni Käufel, 1914 Aphodius reginae Král, 1997 Aphodius spadix Schmidt, 1916 Aphodius swaneticus Reitter, 1892 Aphodius thoracicus Fischer von Waldheim, 1842 † Aphodius aboriginalis Wickham, 1912 † Aphodius anteactus Krell, 2000 † Aphodius bosniaskii Handlirsch, 1907 † Aphodius brevipennis Heer, 1862 † Aphodius charauxi Piton, 1940 † Aphodius cretaceous Nikolajev, 2008 † Aphodius florissantensis Wickham, 1911 † Aphodius granarioides Wickham, 1913 † Aphodius helvolus Statz, 1952 † Aphodius inundatus Wickham, 1914 † Aphodius krantzi Heyden & Heyden, 1866 † Aphodius laminicola Wickham, 1910 † Aphodius mediaevus Wickham, 1914 † Aphodius meyeri Heer, 1847 † Aphodius praeemptor Wickham, 1913 † Aphodius precursor Horn, 1876 † Aphodius schlickumi Statz, 1952 † Aphodius senex Wickham, 1914 † Aphodius shoshonis Wickham, 1912 † Aphodius theobaldi Krell, 2000 † Aphodius vectis Krell, 2019 The species of the following genera were formerly classified in Aphodius: Acrossus Mulsant, 1842 Agoliinus Schmidt, 1913 Alloblackburneus Bordat, 2009 Ballucus Gordon & Skelley, 2007 Blackburneus Schmidt, 1913 Calamosternus Motschulsky, 1859 Caligodorus Gordon & Skelley, 2007 Cephalocyclus Dellacasa et al., 1998 Chilothorax Motschulsky, 1859 Cinacanthus Schmidt, 1913 Coelotrachelus Schmidt, 1913 Colobopterus Mulsant, 1842 Cryptoscatomaseter Gordon & Skelley, 2007 Dellacasiellus Gordon & Skelley, 2007 Dialytodius Gordon & Skelley, 2007 Drepanocanthoides Schmidt, 1913 Eupleurus Mulsant, 1842 Flaviellus Gordon & Skelley, 2007 Geomyphilus Gordon & Skelley, 2007 Haroldiellus Gordon & Skelley, 2007 Hornosus Dellacasa, Dellacasa & Gordon, 2015 Irrasinus Gordon & Skelley, 2007 Labarrus Mulsant & Rey, 1869 Lechorodius Gordon & Skelley, 2007 Liothorax Motschulsky, 1859 Luxolinus Gordon & Skelley,