variation seen in rostral proportions among crocodilians, the latter measurements are probably not an accurate way of estimating body mass and length. Despite this, the species is still one of the largest crocodilians known to have existed, and it may indeed have been the largest gavialoid to have ever existed if a recent revision in the estimated size of the large tomistomine Rhamphosuchus is correct (the genus was once considered to be in length; the new estimate puts it at approximately ). Some skull material also recovered from Peruvian Amazon (Iquitos) in the Pebas Formation of the Middle Miocene, was named as Gryposuchus pachakamue in 2016 by Rodolfo Salas-Gismondi et al. It includes the holotype MUSM 987, a well preserved skull that lacks of temporal and occipital bones; it measures 623.2 millimeters in length, and a series of referred specimens, including possible juveniles. The species was named after the Quechuan word for a primordial god and "storyteller". This new species is characterized by have 22 teeth in the mandible and the maxilla, a snout comparable in relative length to the modern Gavialis gangeticus, and is notable since that its orbits were wider than long and not so upturned as another species of gavialids, including the gryposuchines, which implies that G. pachakamue doesn't had the "telescoped" orbits (protruding eyes) condition fully developed. Since that it species, that inhabited the proto-Amazon fluvial system 13 million years ago, is the oldest record of gavialids in this area and it had a primitive telescoped eyes condition, it shows that the development of such condition was a case of convergent evolution with the species of Gavialis also found in fluvial environments. Indeterminate finds of Gryposuchus were noted from the early Miocene Castillo Formation of Venezuela, middle Miocene Pebas Formation of Peru, middle/late Miocene Tranquitas Formation of Argentina and from the late Miocene formations Urumaco of Venezuela and Solimões in both Brazil and Peru. Additionally, indeterminate finds of gavialoids (all in either coastal or marine sediments) are present in the early Miocene Jimol Formation and the early/middle Miocene Castilletes Formation in Colombia, and from the Oligo-Miocene boundary Pirabas Formation of coastal Brazil. Phylogeny A phylogenetic analysis conducted in a 2007 study found Gryposuchinae to include the genera Aktiogavialis, Gryposuchus, Ikanogavialis, Piscogavialis, and Siquisiquesuchus. Below is a cladogram from the 2007 analysis showing the phylogenetic relationships of gryposuchines among gavialoids: Alternatively, a 2018 tip dating study by