Jennifer Moyle (April 30, 1921 - August 1, 2016) was a British biochemist who helped discover the chemiosmotic mechanism of ATP synthesis. She also conducted research on the properties of purified isocitric enzymes and calcium import in the mitochondria during cellular respiration. Biography Jennifer Moyle was born in Norwich, England and attended Norwich High School for Girls. She was the daughter of S.H. Leonard Moyle and Olive M. Dakin. She had a sister named Vivian, who also studied biochemistry. Moyle began schooling at Norwich High School for Girls in 1926 where she remained until entering Girton College of Cambridge University in 1939. While studying there, she earned a "Title of Bachelor of Arts Degree", the equivalent of modern-day Bachelor of Arts, in 1942. She specialized in Biochemistry, and also attended many lectures on philosophy. In 1955, Moyle began her PhD work at the University of Edinburgh, and was awarded a PhD in zoology in 1958. Moyle conducted a lot of research regarding cellular respiration, oxidative phosphorylation, and properties of purified isocitric enzymes. She was a major contributor to the development of the chemiosmotic theory regarding ATP synthesis. Military service Shortly after earning her degree, Moyle entered the Auxiliary Territorial Service during World War II. She went into military intelligence, where she soon become an intelligence officer in MI8. There, she was promoted to second in command of a section dealing with intelligence obtained from ciphers breaking German codes. After World War II, she continued service for another year helping teach servicemen how to return to civilian life. Research Moyle began her career in biochemistry research in 1964 when she joined a Cambridge biochemistry lab. She began as an assistant to Marjory Stephenson, but after a few years Stevenson introduced Moyle to Peter D. Mitchell, with whom Moyle worked for the majority of her research career. Jennifer's main research focus was cellular respiration. for a short period, Moyle worked with Malcolm Dixon, and the two focused their work on the purification of isocitric enzymes. Purified isocitric enzymes Jennifer Moyle published an article on the properties of purified isocitric enzymes in August 1956 with Malcolm Dixon. The article describes the chemical and physical properties of isocitric enzymes, various reactions they are involved in, causes for their inactivation, and a hypothesis for the mechanism of reaction. Moyle and Dixon authored another paper regarding the isocitric enzyme Triphosphopyridine Nucleotide-linked isoCitric Dehydrogenase-Oxalosuccinic Carboxylase. The paper