elsewhere in the region. The missionary R. W. Felkin, who had met the ruler, remarked that Mutesa "had lost the pure Hamitic features through admixture of Negro blood, but still retained sufficient characteristics to prevent all doubt as to his origin". Thus, Keane would suggest that the original Hamitic migrants to the Great Lakes had "gradually blended with the aborigines in a new and superior nationality of Bantu speech". Speke believed that his explorations uncovered the link between "civilized" North Africa and "primitive" central Africa. Describing the Ugandan Kingdom of Buganda, he argued that its "barbaric civilization" had arisen from a nomadic pastoralist race who had migrated from the north and was related to the Hamitic Oromo (Galla) of Ethiopia. In his Theory of Conquest of Inferior by Superior Races (1863), Speke would also attempt to outline how the Empire of Kitara in the African Great Lakes region may have been established by a Hamitic founding dynasty. These ideas, under the rubric of science, provided the basis for some Europeans asserting that the Tutsi were superior to the Hutu. In spite of both groups being Bantu-speaking, Speke thought that the Tutsi had experienced some "Hamitic" influence, partly based on their facial features being comparatively more narrow than those of the Hutu. Later writers followed Speke in arguing that the Tutsis had originally migrated into the lacustrine region as pastoralists and had established themselves as the dominant group, having lost their language as they assimilated to Bantu culture. Seligman and other early scholars believed that, in the African Great Lakes and parts of Central Africa, invading Hamites from North Africa and the Horn of Africa had mixed with local "Negro" women to produce several hybrid "Hamiticised Negro" populations. The "Hamiticised Negroes" were divided into three groups according to language and degree of Hamitic influence: the "Negro-Hamites" or "Half-Hamites" (such as the Maasai, Nandi and Turkana), the Nilotes (such as the Shilluk and Nuer), and the Bantus (such as the Hima and Tutsi). Seligman would explain this Hamitic influence through both demic diffusion and cultural transmission: European colonial powers in Africa were influenced by the Hamitic hypothesis in their policies during the twentieth century. For instance, in Rwanda, German and Belgian officials in the colonial period displayed preferential attitudes toward the Tutsis over the Hutu. Some scholars argued that this bias was a significant factor that contributed to the 1994 Rwandan genocide