with Tupuxuara longicristatus, and he also indicated that the paratype of S. brasiliensis was likely referable to T. longicristatus. Kellner (1990) renamed S. araripensis to Anhanguera araripensis, followed by Wang et al. (2008), though Veldmeijer (2003) included it in Coloborhynchus. Recent study, however, considers S. araripensis dubious. Classification De Buisonjé first assigned Santanadactylus to Criorhynchidae (=Ornithocheiridae). Wellnhofer (1991) considered it a member of the Ornithocheiridae based on the structure of the humerus, but noted that the elongation of the paratype neck vertebrae distinguished it from other ornithocheirids. Bennett (1989) assigned S. brasilensis (holotype only) to Pteranodontidae. Kellner (1990), concluding that he could find but a single autapomorphy for Santanadactylus brasilensis, the straight ventral margin of the proximal part of the deltopectoral crest, assigned the genus to Pterodactyloidea incertae sedis, but later (Kellner and Tomida 2000) assigned it to Pteranodontoidea. S. pricei according to Kellner belonged to a clade descending from the last common ancestor of Istiodactylus and the Anhangueridae. The same was in his analysis true for Araripesaurus, a genus of which he had previously thought S. pricei was a junior synonym. Paleobiology Santanadactylus is regarded as a large pterosaur, Wellnhofer for the various species indicating a wingspan of 2.9–5.7 meters (9.5–18.7 ft). De Buisonjé thought Santanadactylus brasilensis had a wingspan of . It may have been adapted for gliding over flapping flight. See also List of pterosaur genera Timeline of pterosaur research Pterosaur size References External links Santanadactylus in The Pterosauria. Accessed 2007–02–09 Re: Pterosaur Help, a posting from George Olshevsky on the Dinosaur Mailing List, which, although incomplete, should give some idea as to the complexity of the taxonomy here. Accessed 2007–02–09 Pteranodontoids Early Cretaceous pterosaurs of South America Cretaceous Brazil Fossils of Brazil Albian life Fossil taxa described in 1980