of uncertain or fairly basal position. Fossils Numerous fossil Hemerobiidae have been described, some from the still-living genera, others from genera that are entirely extinct today. While most have been found in Eocene to Miocene rocks or amber, Promegalomus is known from the Jurassic. It was formerly considered to constitute a distinct family Promegalomidae, but is currently recognized as a very basal member of the Hemerobiidae. The Cretaceous Mesohemerobius was formerly considered a brown lacewing, but is today rather placed as incertae sedis in the Neuroptera; it might be a member of the Hemerobioidea but not even that is certain. Notable fossil Hemerobiidae genera are: Bothromicromus Scudder, 1878 (Eocene/Oligocene; Quesnel, British Columbia) Brasilopsychopsis Crato Formation Brazil, Early Cretaceous (Aptian) Cratopsychopsis Crato Formation Brazil, Aptian Cretomerobius Ponomarenko, 1992 Dzun-Bain Formation, Mongolia, Aptian Hemeroberotha Makarkin & Gröhn, 2020 Burmese amber, Myanmar, Late Cretaceous (Cenomanian) Mucropalpus Pictet, 1856 (Eocene; Baltic amber) Plesiorobius Klimaszewski and Kevan 1986 Late Cretaceous (Taimyr amber, Russia, Santonian Canadian amber, Campanian Ola Formation, Russia, Campanian) Prochlanius Kruger, 1923 (Eocene; Baltic amber) Promegalomus Panfilov, 1980 Karabastau Formation, Kazakhstan, Middle/Late Jurassic (Callovian/Oxfordian) Prophlebonema Kruger, 1923 (Eocene; Baltic amber) Prospadobius Kruger, 1923 (Eocene; Baltic amber) Purbemerobius Jepson et al. 2012 Durlston Formation, United Kingdom, Early Cretaceous (Berriasian) The extinct genus Hemerobites was originally described in 1813 from a specimen preserved in Baltic amber as a hemerobiid. However further study has resulted in its synonymy with the genus Eutermes and placement in the termite subfamily Termitina. The species Wesmaelius mathewesi, was described in 2003 from a solitary Eocene fossil found near Quesnel, British Columbia, and placed into the extant genus Wesmaelius. At that time it was the oldest Hemerobiinae species described. Another species from the same genus, Wesmaelius makarkini was found in Garang Formation of Zeku County, Qinghai Province, China, in 2018. The finding is from Lower Miocene. Some additional brown lacewing larvae have been found as fossils, but it has been impossible to determine their generic or subfamilial association. References (1986): Collins Guide to the Insects of Britain and Western Europe. External links BioLib Taxonomic tree of Hemerobiidae (incomplete) Brown lacewings of Florida on the UF / IFAS Featured Creatures Web site Hemerobiiformia Neuroptera Neuroptera families Articles containing video clips Extant Jurassic first appearances