Jonathan Lindsay Penner (born March 5, 1962) is an American actor, screenwriter, television personality, and film producer, known for producing and starring in the film The Last Supper, as well as acting in the television series Rude Awakening and The Naked Truth. He is also known for his multiple appearances on the American competitive reality series Survivor. Life and career Penner was born in New York City. He has had roles in television series Grapevine, The Naked Truth and Rude Awakening, and guest-starred on Seinfeld, CSI, CSI: NY, The Tick, Melrose Place, LateLine, Arrested Development, and The Nanny, where he appeared in two episodes as Danny Imperiali, whose firing and dumping of fiancée Fran Fine kicked off the entire series. Penner has appeared in many films, with featured roles in Anarchy TV, Down Periscope, and The Last Supper, the latter of which he also co-produced, and which was directed by his wife, Stacy Title. He had a small role in Dragon: The Bruce Lee Story (1993). Penner was nominated for Best Live Action Short Film at the Academy Awards in 1994 for Down on the Waterfront, which he co-produced, co-wrote, and starred in, along with Jason Alexander and Edward Asner. He co-wrote the script for the 2003 The WB-produced re-imagining of The Lone Ranger, and later the 2017 horror film The Bye Bye Man. He is developing a King Kong television series for MarVista Entertainment and IM Global Television. Penner resides in Los Angeles, California, with his two children: Ava, born in 1999, and Cooper, born in 1995. He was married to director Stacy Title from 1991 until her death in 2021; he was introduced to Stacy by his friend Jason Alexander, who is married to Stacy's cousin, Daena Title. Filmography Survivor In addition to his acting and writing credits, Penner is perhaps best known as a three-time contestant on Survivor. Cook Islands Penner was a contestant on Survivor: Cook Islands in 2006. He was one of the original five members of the Rarotonga tribe (Raro). He quickly became notorious among the cast for stealing another tribe's chicken on the first day. In episode three, the four tribes evenly divided into two new mixed tribes, Aitutaki and Rarotonga. Penner found himself on Aitutaki (Aitu). While a member of Aitu, he became part of an alliance that consisted of Yul Kwon, Candice Woodcock, Becky Lee and Sundra Oakley. This alliance lasted