on 6 August, by French forces from Dahomey to the east and on 9 August by British forces from Gold Coast to the west. German colonial forces withdrew from the capital Lomé and the coastal province and then fought delaying actions on the route north to Kamina, where a new wireless station linked Berlin to Togoland, the Atlantic and South America. The main British and French force from the neighbouring colonies of Gold Coast and Dahomey advanced from the coast up the road and railway, as smaller forces converged on Kamina from the north. The German defenders were able to delay the invaders for several days at the battles of Bafilo, Agbeluvhoe and Chra but surrendered the colony on 26 August 1914. In 1916, Togoland was partitioned by the victors and in July 1922, British Togoland and French Togoland were created as League of Nations mandates. The French acquisition consisted of of the colony, including the coast. The British received the smaller, less populated and less developed portion of Togoland to the west. The surrender of Togoland was the beginning of the end for the German colonial empire in Africa. Bussa uprising, 1915 In the Colony and Protectorate of Nigeria (now Nigeria), the British policy of indirect rule through local proxies was extended after the outbreak of the First World War, when British colonial officers and troops were withdrawn for war service. The British became more dependent on local emirs but in Bussa, the re-organisation of local government in 1912 overthrew the authority of the traditional ruler. The hereditary Emir of Bussa, Kitoro Gani was moved aside and the Borgu Emirate was divided, each area ruled by a (native administration). In June 1915, about 600 rebels, armed with bows and arrows occupied Bussa, captured and killed half of the new native administration; the survivors fled the district. The rebellion caused panic because the British authorities were so short of troops. A small force from the West African Frontier Force (WAFF) and the Nigeria police moved into Bussa and skirmished with the rebels. No soldiers were killed and only 150 shots were fired. Sabukki, one of the ringleaders fled to nearby French Dahomey and the rebellion was suppressed. The war in Nigeria played a role in British politics during the war. At the beginning of the war, the British government seized foreign expatriate firms in the lucrative palm oil trade owned