only from juvenile specimens). In 2018, Moore and colleagues redescribed Bellusaurus and provided several arguments to refute this notion. First, they noted that the two were not actually contemporaries; the holotype of Klamelisaurus originates from slightly older strata (rock layers). They also listed twenty-four characteristics in the vertebrae, coracoid, and humerus that differentiated the two genera. In their 2020 redescription of Klamelisaurus, they added four characteristics to this list, and also noted that Bellusaurus did not possess any of the unique features of Klamelisaurus. Most of the characteristics on their list pertain to differences in pneumatic features in the vertebrae of Klamelisaurus and Bellusaurus that are not readily explainable by differences in age. In sauropods, vertebral air spaces typically become more extensive and invade more of the vertebrae with age. Bellusaurus possessed procamerate to weakly camerate air spaces (i.e., its air spaces were deep, leaving only a thin layer of bone at the midline, and were barely enclosed by bone) in its cervicals. Meanwhile, based on comparisons with its relatives, Moore and colleagues inferred that Klamelisaurus likely had camerate air spaces (i.e., enclosed by bone) in its cervicals. Based on the cervicals alone, they could not contradict the hypothesis that juvenile Bellusaurus could have grown into an adult with camellate air spaces in the cervicals, as is the case with Barosaurus. However, they noted that Bellusaurus possessed several pneumatic features in the dorsal neural spines and that Klamelisaurus lacked, which runs contrary to the expected pattern for euhelopodids and mamenchisaurids. Specifically, in the dorsals of Bellusaurus, the PCDLs (posterior centrodiapophyseal laminae) below the diapophyses bifurcated at the bottom; the CPOLs (centropostzygapophyseal laminae) at the rear of the centra had a sharp-edged branch situated in a relatively deep depression; and the diapophyses bore rimmed trough-like depressions. Considering their temporal and anatomical differences, Moore and colleagues thus considered Klamelisaurus and Bellusaurus to be readily distinguishable. However, they also noted two characteristics on their list which may have varied with age: the bifurcated neural spines, and the presence of wing-shaped processes that projected further outwards than the postzygapophyses on the rear dorsals. Palaeoenvironment The holotype of Klamelisaurus originated from a rock layer that was described as being "gray-brown, purple-red sandy mudstone" by Zhao in 1993. This layer was situated at the top of what Zhao called the "Wucaiwan Formation", but a lack of differences in rock layers has led it to become subsumed