moss, floods during rain. The eggs hatch and the tadpoles develop in the water. If no sufficient rain happens soon after laying the eggs can remain unhatched for many weeks, with the tadpoles developing inside. The nest must stay wet so the eggs remain moist, but not too wet or they will be washed away. Female frogs that mate with more males, thus depositing eggs in more nest sites, are more likely to have some young survive. Calling behavior This species attracts mates and establishes its territory by displaying certain calls during the breeding season. This calling behavior uses acoustic signals that are modulated by the presence of other males and females in the area. There are two characteristic calls of this species. The first one is the “advertisement call” which is marked by a short two part sequence that sounds at a higher relative frequency. The second call is the “territorial call” which is distinguished by its longer one part call that sounds at a lower overall frequency. The frequency of these calls is greatly affected by the presence of other members of the species in the area. In response to increased presence of males the call rate increased by twofold, and in response to the increased presence of females the call rate increased by threefold. Additionally, this species increases the frequency of territorial calls in the presence of males and increases advertisement calls in the presence of females. Thus, each of these calls has a specific effect on female choice and male competition that determines the overall mating success of each individual. Males only reach the maximum level of advertising calls in the presence of females. If there are only males present, the males will choose to wait until there is a possibility they could mate. Also, females have a bias towards males with more complex calls produced at a higher rate. It has been thought that better calling patterns are an indication of the quality of the nest site which may be why they are so important in mating success. Overall, it is evident that calling behavior is significantly affected by competition amongst males and the possibility of mating with females, and calling behavior clearly affects females in their decisions to mate and reproduce. Chemosignals For this species, calling behavior during mating season can have significant costs for the survival of the frog. Time spent calling decreases