into the Shishugou Formation as the "lower beds" or the Wucaiwan Member. Using stratigraphic correlation, this rock layer was found to lie below a tuff from the Shishugou Formation at the Wucaiwan locality, which has been dated radiometrically to 162.2 ± 0.2 Ma, or the Oxfordian age of the Jurassic period. Based on this, Moore and colleagues considered Klamelisaurus to have originated from the late Callovian age. During the Callovian, the climate of the Shishugou Formation is considered to have been mesic (moderately and seasonally wet), with the environment at Wucaiwan having been an alluvial plain or marsh. Juliane Hinz and colleagues in 2010 reconstructed a petrified forest preserved in overlying Oxfordian rocks, located north of Jiangjunmiao. It would have consisted of Araucaria trees, with the undergrowth being occupied by Coniopteris tree ferns, Anglopteris and Osmunda ferns, Equisetites horsetails, and Elatocladus shrubs. Three theropod dinosaurs have been discovered near Jiangjunmiao: Monolophosaurus, considered to have been no younger than the late Callovian and thus closest temporally to Klamelisaurus; Aorun, from layers in the "middle beds" that have been re-dated from the Callovian to the Oxfordian; and Sinraptor, from the Oxfordian "upper beds". The crocodylomorphs Sunosuchus and Junggarsuchus are known from other localities in the "lower beds". Meanwhile, the sauropods Bellusaurus, Mamenchisaurus sinocanadorum, and Tienshanosaurus are known from the "upper beds", above the level of the tuff at 162.2 Ma, and thus were not contemporaries of Klamelisaurus. Asides from these sauropods, Aorun, and Sinraptor, the Oxfordian portion of the Shishugou Formation preserves a diverse dinosaur fauna that also includes the theropods Haplocheirus, Shishugounykus, Zuolong, Guanlong, and Limusaurus; the ornithopod Gongbusaurus; the stegosaur Jiangjunosaurus; and the marginocephalians Yinlong and Hualianceratops. References Sauropods Middle Jurassic dinosaurs of Asia Jurassic China Biota of Xinjiang Paleontology in Xinjiang Taxa named by Zhao Xijin