Reiche, 1841 Parachorius Versicorpus Deschodt Davis and Scholtz, 2011 Onthophagini - about 40 genera including: Hyalonthophagus Onthophagus Proagoderus Predominantly Old World (New World presence likely represents rather recent dispersal) Oniticellini (incl. Eurysternini) - 16 genera including: Euoniticellus Janssens, 1953 Oniticellus Serville, 1825 Eurysternus Sisyphini - three genera (sometimes placed in the Scarabaeini): Nesosisyphus Neosisyphus Sisyphus Old World Gymnopleurini - four genera (sometimes placed in the Scarabaeini): Allogymnopleurus Garreta Janssens, 1940 Gymnopleurus Illiger, 1803 Paragymnopleurus Scarabaeini - selected genera: Scarabaeus (Kheper is a subgenus) Onitini - selected genera: Bubas Mulsant, 1842 Cheironitis Lansberge, 1875 Onitis Fabricius, 1798 New World Eucraniini - four genera: Anomiopsoides Ennearabdus Eucranium Glyphoderus Phanaeini - 12 genera including: Oxysternon Phanaeus References External links Subfamily Scarabaeinae: photographs of representatives of the tribes Extant Eocene first appearances