named by Marsh, P. nanus and P. altus, as different growth stages of Astrodon johnstoni. However, other authors did not find the argument in favor of the synonymization of Astrodon and Pleurocoelus so convincing. According to Peter Rose (2007) it has not been demonstrated that either the teeth of Astrodon johnstoni or those attributed to Pleurocoelus are "morphologically diagnostic among titanosauriforms", which limits their utility when it comes to distinguishing them from the teeth of other taxa. The type series of Pleurocoelus nanus and P. altus (four vertebrae and two hindlimbs bones, respectively) cannot be directly compared to the teeth from the type series of Astrodon, so any comparison has to be conducted based on the referred specimens of Pleurocoelus. These, however, are all isolated bones from the Arundel Formation, which themselves were referred to Pleurocoelus only based "on proximity of the localities and the size of the bones". Rose concludes that, as Astrodon is not based on the diagnostic material, "new discoveries should not be aligned with that genus" and that "the argument to synonymize the two taxa, Astrodon and Pleurocoelus, seems unfounded". The type material of Pleurocoelus may not be diagnostic as well, according to the author. A similar argument was made by Michael D. d'Emic (2013). The author did not find any diagnostic features of the type material of Astrodon johnstoni, Pleurocoelus nanus and P. altus and considered the three taxa to be nomina dubia; according to the author there is no direct evidence that any sauropod bones from the Arundel Formation other than their type series can be referred to these taxa. D’Emic also stated that the exact provenance of the bones from the type series of Pleurocoelus nanus is uncertain and thus "these bones could represent a chimera of individuals or taxa". The author also commented on the diagnosis of Astrodon johnstoni proposed by Carpenter and Tidwell (which was based on all of the sauropod material from the Arundel Formation, not only on the teeth from the type series); he claimed that most of the supposed autapomorphies of this taxon "are indistinguishable compared to other sauropods such as Camarasaurus (...) and/or are related to the juvenile nature of the material". Paleoecology Habitat The Arundel Formation of Maryland has been dated through palynomorphs to the Albian stage of the Early Cretaceous period, about 112 million years ago. This formation is part of the Potomac Group that