from any shark, though it also reflects the concentration of force at the tooth tip. Dense aggregations of young sharks, forming in response to feeding opportunities, have been documented in the Indian Ocean. The known diet of the dusky shark encompasses pelagic fishes, including herring and anchovies, tuna and mackerel, billfish, jacks, needlefish and flyingfish, threadfins, hairtails, lancetfish, and lanternfish; demersal fishes, including mullets, porgies, grunts, and flatheads, eels, lizardfish, cusk eels, gurnards, and flatfish; reef fishes, including barracudas, goatfish, spadefish, groupers, scorpionfish, and porcupinefish; cartilaginous fishes, including dogfish, sawsharks, angel sharks, catsharks, thresher sharks, smoothhounds, smaller requiem sharks, sawfish, guitarfish, skates, stingrays, and butterfly rays; and invertebrates, including gastropods, cephalopods, decapod crustaceans, barnacles, and sea stars. Very rarely, the largest dusky sharks may also consume sea turtles, marine mammals (mainly as carrion), and human refuse. In the northwestern Atlantic, around 60% of the dusky shark's diet consists of bony fishes, from over ten families with bluefish (Pomatomus saltatrix) and summer flounder (Paralichthys dentatus) being especially important. Cartilaginous fishes, mainly skates and their egg cases, are the second-most important dietary component, while the lady crab (Ovalipes ocellatus) is also a relatively significant food source. In South African and Australian waters, bony fishes are again the most important prey type. Newborn and juvenile sharks subsist mainly on small pelagic prey such as sardines and squid; older sharks over long broaden their diets to include larger bony and cartilaginous fishes. The run of the southern African pilchard (Sardinops sagax), occurring off the eastern coast of South Africa every winter, is attended by medium and large-sized dusky sharks. Pregnant and post-partum females do not join, possibly because the energy cost of gestation leaves them unable to pursue such swift prey. One South African study reported that 0.2% of the sharks examined had preyed upon bottlenose dolphins (Tursiops truncatus). Life history Like other requiem sharks, the dusky shark is viviparous: the developing embryos are initially nourished by a yolk sac, which is converted into a placental connection to the mother once the yolk supply is exhausted. Mating occurs during spring in the northwestern Atlantic, while there appears to be no reproductive seasonality in other regions such as off South Africa. Females are capable of storing masses of sperm, possibly from multiple males, for months to years within their nidamental glands (an organ that secretes egg cases). This would be advantageous given the sharks' itinerant