the head from an early age. The Cape sparrow's calls are chirps similar to those of the house sparrow, but much more musical and mellow. The basic call is used in flight and while perching socially and transcribed as chissip, chirrup, , or . A loud, distinctive call used by the male to advertise nest ownership can be written as tweeng or twileeng; this call can be extended into a jerky and repetitive song, chip cheerup, chip cheerup. Taxonomy The Cape sparrow was first taxonomically described by Philipp Ludwig Statius Müller in 1776, as Loxia melanura. Some other earlier biologists described the Cape sparrow in Loxia or Fringilla, but it has otherwise been regarded as a member of the genus Passer along with the house sparrow and other sparrows of the Old World. Within this genus, morphological comparisons and geography were insufficient to elucidate to which species the Cape sparrow is most closely related. Mitochondrial DNA phylogenies have strongly suggested that the Cape sparrow is among the most basal members of its genus, having diverged from the rest of the genus during the late Miocene, over 5 million years ago. It is genetically closest to the southern grey-headed sparrow and the other grey-headed sparrows of Africa and the saxaul sparrow of Central Asia, so these species may be sibling species of the Cape sparrow or similarly early offshoots. The Cape sparrow's specific epithet comes from the Greek μέλας (melas, "black") and ουρά (oura, "tail"), while the name of the genus Passer comes from a Latin word for small birds. The Cape sparrow has three subspecies. The nominate subspecies Passer melanurus melanurus is found in western South Africa, east to the western part of Free State. The subspecies vicinus, which is sometimes included in subspecies melanurus, occurs from Free State east to Eastern Cape and Lesotho. The subspecies damarensis ranges from the extreme southern coastal areas of Angola into Namibia, Botswana and southern Zimbabwe, as well as northern South Africa. Distribution and habitat The Cape sparrow inhabits southern Africa south of Angola and as far east as Eswatini. The northernmost point in its range is Benguela in Angola, and it is found in the coastal and central parts of Namibia, except for the driest parts of the Namib Desert. It occurs in all of South Africa except the farthest east, in southern Botswana and spottily in the Kalahari Basin of central