Diodorus, a paratypothoracisine aetosaur, and procolophonid parareptiles, as well as the stahleckeriid dicynodont Moghreberia, a synapsid. Temnospondyl amphibians are represented by several genera of metoposauroids, including the metoposaurids Arganasaurus and Dutuitosaurus, and the latiscopid Almasaurus. Fish are also known from the T5 member, including various ray-finned actinopterygians such as the locally endemic Dipteronotus gibbosus and Mauritanichthys, as well as other perleidiform and redfieldiiform fishes, alongside lobe-finned actinistians and lungfish such as Asiatoceratodus. The T5 member is composed of cyclical layers of bioturbated mudstone and sandstone deposited in a broad, semi-humid basin. It has been interpreted as a system of brackish permanent lakes and sandbars, or alternatively sandy meandering rivers on a muddy floodplain. The fluvial or lacustrine sediments of the T5 member contrast with the playa sediments of the preceding T4 member, suggesting that it was deposited during an interval of increased rainfall. Numerous tracks and trackways from various animals are preserved, typically those of animals known from fossil remains such as phytosaurs, pseudosuchians, dinosauromorphs and basal archosauromorphs. The tracks also appear to indicate the presence of large to very-large dinosauromorphs or paracrocodylomorphs that are currently not yet known from skeletal remains. Additional traces mark the presence of burrowing invertebrates, bivalves, and clam shrimps. Makay Formation, Madagascar In Madagascar, Azendohsaurus co-existed with the hyperodapedontine rhynchosaur Isalorhynchus, the herbivorous traversodontid cynodonts Dadadon and Menadon, and the predatory chiniquodontid cynodont Chiniquodon kalanoro, as well as an undescribed kannemeyeriiform dicynodont, a sphenodontian reptile, a procolophonid parareptile, the diminutive lagerpetid Kongonaphon, various other undescribed dinosauromorphs, and an "enigmatic archosaur" of uncertain classification. The faunal composition of the Isalo II is believed to represent a Middle Triassic Ladinian aged assemblage, existing prior to the appearance of dinosaurs and associated Late Triassic faunas, particularly aetosaurs and phytosaurs that are absent from the formation, and also inferred from the dominance of traversodonts in the fauna. However, this age assessment remains uncertain, and the formation is possibly from the younger early Late Triassic during the Carnian, as has been proposed for the T5 member of the Argana Formation. Fossils of A. madagaskarensis have been exclusively recovered from a deposit of fine grained red mudstone, while other fossil bearing localities in the formation consist of medium grained channel sands, possibly reflecting a habitat preference in the ecosystem distinct from other animals or unique behavioural trait. The absence of any other species in the bone bed may also support this.