related to them. This was based on the characteristics of its vertebrae, as the cranial remains had not yet been found. In 1994 these cranial remains would be found - two maxilla - and these would be used by the original authors to establish it as a species of hadrosaur. Casanovas-Cladellas et al. revised their position for a final time in 1999, when they published a paper arguing for a position as a primitive member of the hadrosaur subfamily Lambeosaurinae - this made it the first such species known from the continent of Europe. Characters of the skeleton supporting this viewpoint were the truncated and rounded anatomy of the articulation of the maxilla to the jugal, the truncated nature of the back of the maxilla itself, the ventral deflection of the front of the dentary (thought the only known dentary was later referred to a new species, Koutalisaurus, and later declared that of indeterminate lambeosaurine), its tall neural spines, and of the humerus being distally projected. The proposal that P. isonensis was the first known European lambeosaurine was soon challenged, however, in 2001 by Jason Head, in a study re-evaluating the status of another species, Eolambia caroljonesa, as a primitive lambeosaur. Both Pararhabdodon and Eolambia were found to instead be more primitive hadrosauroids more basal than the split between Lambeosaurinae and "Hadrosaurinae" (later renamed to Saurolophinae). In regards to Pararhabdodon, this was based on a rebuttal of arguments put forward in the 1999 study; the present anatomy maxilla-jugal articulation was not considered a synapomorphy in Lambeosaurinae, merely the presence of this in the jugal itself, unknown in the species, and the angular deltopectoral crest is a trait present even in primitive iguanodontians. Additionally, its tooth count was lower than those of known lambeosaurs. In the 2006 re-evaluation of the genus by Albert Prieto-Márquez and colleagues, it was included in a phylogenetic analysis for the first time. This found it to be a non-hadrosaurid hadrosauroid, a similar position as had been argued by Head. The cladogram of Prieto-Márquez et al. (2006) is seen below on the left: Prieto-Márquez would return to the issue in 2009 along with Jonathan R. Wagner. They once again turned to the articulation between the maxilla and the jugal, finding this to link Pararhabdodon to the Asian lambeosaurine, Tsintaosaurus spinorhinus. In the 2006 phylogenetic analysis, P. isonenis had been scored as having an ancestral hadrosauroid condition for