waves of admixture, which had more impact on higher-caste and northern Indians and took place more recently. This may be explained by "additional gene flow", related to the spread of languages: Palanichamy et al. (2015), elaborating on Kivisild et al. (1999) conclude that "A large proportion of the west Eurasian mtDNA haplogroups observed among the higher-ranked caste groups, their phylogenetic affinity and age estimate indicate recent Indo-Aryan migration to India from west Asia. According to Palanichamy et al. (2015), "the west Eurasian admixture was restricted to caste rank. It is likely that Indo-Aryan migration has influenced the social stratification in the pre-existing populations and helped in building the Hindu caste system, but it should not be inferred that the contemporary Indian caste groups have directly descended from Indo-Aryan immigrants. Jones et al. (2015) state that Caucasus hunter gatherer(CHG) was "a major contributor to the Ancestral North Indian component". According to Jones et al. (2015), it "may be linked with the spread of Indo-European languages", but they also note that "earlier movements associated with other developments such as that of cereal farming and herding are also plausible". note that the ANI are inseparable from Central-South Asian populations in present-day Pakistan. They hypothesise that "the root of ANI is in Central Asia". According to Lazaridis et al. (2016) ANI "can be modelled as a mix of ancestry related to both early farmers of western Iran and to people of the Bronze Age Eurasian steppe". Silva et al. (2017) state that "the recently refined Y-chromosome tree strongly suggests that R1a is indeed a highly plausible marker for the long-contested Bronze Age spread of Indo-Aryan speakers into South Asia." Silva et al. (2017) further notes "they likely spread from a single Central Asian source pool, there do seem to be at least three and probably more R1a founder clades within the Subcontinent, consistent with multiple waves of arrival." conclude that pastoralists spread southwards from the Eurasian steppe during the period 2300–1500 BCE. These pastoralists during the 2nd millennium BCE, who were likely associated with Indo-European languages, presumably mixed with the descendants of the Indus Valley Civilisation, who in turn were a mix of Iranian agriculturalists and South Asian hunter-gatherers forming "the single most important source of ancestry in South Asia." Zerjal et al. (2002) argue that "multiple recent events" may have reshaped India's genetic landscape. Origins of R1a-Z93 Ornella Semino et al. (2000) proposed