Some forms of warning coloration provide this distance dependent camouflage by having an effective pattern and colour combination that do not allow for easy detection by a predator from a distance, but are warning-like from a close proximity, allowing for an advantageous balance between camouflage and aposematism. Warning coloration evolves in response to background, light conditions, and predator vision. Visible signals may be accompanied by odours, sounds or behaviour to provide a multi-modal signal which is more effectively detected by predators. Unpalatability, broadly understood, can be created in a variety of ways. Some insects such as the ladybird or tiger moth contain bitter-tasting chemicals, while the skunk produces a noxious odour, and the poison glands of the poison dart frog, the sting of a velvet ant or neurotoxin in a black widow spider make them dangerous or painful to attack. Tiger moths advertise their unpalatability by either producing ultrasonic noises which warn bats to avoid them, or by warning postures which expose brightly coloured body parts (see Unkenreflex), or exposing eyespots. Velvet ants (actually parasitic wasps) such as Dasymutilla occidentalis both have bright colours and produce audible noises when grabbed (via stridulation), which serve to reinforce the warning. Among mammals, predators can be dissuaded when a smaller animal is aggressive and able to defend itself, as for example in honey badgers. Prevalence In terrestrial ecosystems Aposematism is widespread in insects, but less so in vertebrates, being mostly confined to a smaller number of reptile, amphibian, and fish species, and some foul-smelling or aggressive mammals. Pitohuis, red and black birds whose toxic feathers and skin apparently comes from the poisonous beetles they ingest, could be included. It has been proposed that aposematism played a role in human evolution, body odour carrying a warning to predators of large hominins able to defend themselves with weapons. Perhaps the most numerous aposematic vertebrates are the poison dart frogs (family: Dendrobatidae). These neotropical anuran amphibians exhibit a wide spectrum of coloration and toxicity. Some species in this poison frog family (particularly Dendrobates, Epipedobates, and Phyllobates) are conspicuously coloured and sequester one of the most toxic alkaloids among all living species. Within the same family, there are also cryptic frogs (such as Colostethus and Mannophryne) that lack these toxic alkaloids. Although these frogs display an extensive array of coloration and toxicity, there is very little genetic difference between the species. Evolution of their conspicuous coloration is