selection at work. Darwin preferred to leave the evolutionary implications hanging. Korn points out that the book has never been out of print since 1872, calling into question Ekman's talk of "Darwin's lost masterpiece". The "Editor's notes" at the "Mead Project source page" on the book comment that Publication Darwin concluded work on the book with a sense of relief. The proofs, tackled by his daughter Henrietta ("Ettie") and son Leo, required a major revision which made Darwin "sick of the subject and myself, and the world". The Expression was published by John Murray on 26 November 1872. It quickly sold around 7,000 copies and was widely praised as a charming and accessible introduction to Darwin's evolutionary theories. A second edition was published by Darwin's son in 1890, without several revisions suggested by Darwin; these were not published until the third edition of 1999 (edited by Paul Ekman). Influence Published as a sequel to The Descent of Man, The Expression was assured of a wide readership in mid-Victorian England. However, the early death of George Romanes (1848–1894) robbed Darwin of a powerful advocate in the field of comparative psychology and his impact on academic psychology was muted, partly because of Wilhelm Wundt's dimensional approach to the emotions and the widespread influence of the behaviorist school during the twentieth century. The generous style of biological illustration was followed in work on animal locomotion by photographer Eadweard Muybridge (1830–1904) (leading to cinematography), and by the Scottish naturalist James Bell Pettigrew (1832–1908); in the extensively (and controversially) illustrated works of the evolutionary biologist Ernst Haeckel; and – to a lesser extent – in D'Arcy Thompson's On Growth and Form (1917). Darwin's ideas were followed up in William James' What Is An Emotion ? (1884); and, in the James-Lange theory of emotions, James develops Darwin's emphasis on the physical aspects, including the visceral (autonomically mediated) components of emotion. In Walter Cannon's Bodily Changes in Pain, Hunger, Fear and Rage (1915), Cannon introduces the famous phrase fight or flight response, formulating emotions in terms of strategies for interpersonal behaviour and amplified in groups or crowds (herd behavior). More recent psychological theories of emotion have been set out in the Papez-Maclean hypothesis, the Two factor theory of emotion (Schachter and Singer) and the Theory of constructed emotion. On 24 January 1895, James Crichton-Browne delivered a notable lecture in Dumfries, Scotland On Emotional Expression, presenting some of