Nikolaj Vladimirovich Timofeev-Resovskij (also Timofeyeff-Ressovsky; ; – 28 March 1981) was a Soviet biologist. He conducted research in radiation genetics, experimental population genetics, and microevolution. His life was highlighted by scientific achievements in the face of severe personal hardship, including his imprisonment and working in secret scientific facilities of Soviet Gulag. Timofeev-Ressovsky was a descendant of the old Russian school of scientists, characterised by broad naturalistic views on the world, simultaneously combined with exact analysis of causes and consequences and establishment of elementary phenomena. He widely collaborated with physicists. Known for his influential personality, he was a talented story-teller and teacher. Education Nikolaj Vladimirovich Timofeev-Resovski, began his university education from 1916 to 1917 at the Moscow City People's University named after A. L. Shanyavskij. From 1917 to 1922, he studied at the First Moscow State University. The First World War and the consequences of the Russian Revolution of 1917 interrupted his education for periods of time. At the outbreak of the Russian Civil War, Timofeev-Resovskij was a follower of the anarchist Peter Kropotkin. In 1918, he volunteered to serve in a small anarchist cavalry unit, which was part of the Green army, i.e., they were neither supporters of the Bolshevik Red army nor the White army of General Anton Ivanovich Denikin. Eventually, in 1919, the anarchists joined the Red army. As a private in the 12th Red Army, Timofeev-Resovskij took part in the last battle of the Civil War in the Crimea on the Polish front. A paper trail of his education was swept away in the chaos of war. Timofeev-Resovskij abandoned his education without a scientific degree; he left formal education with only a gold medal. Even though he eventually became a recognized world-class scientist, he would later, as a requirement for employment, be required to complete his doctorate. He defended his doctoral dissertation in 1963 in Sverdlovsk, and he was awarded a Doctor of Science diploma in 1964. Career In Russia During the period 1920 to 1925, Timofeev-Resovskij was an instructor and researcher in Moscow. It was in this period that he met and married Elena Aleksandrovana Fidler (1898 – 1973), who was a student of the cytologist and biologist Nikolaj Konstantinovich Kol’tsov in Moscow; Elena became not only Timofeev-Resovskij's lifelong devoted companion but also his lifelong partner in research. Their eldest son, Dmitrij, was born on 11 September 1923. From 1920 to 1925, Timofeev-Resovskij was a biology