Sir Paul Maxime Nurse (born 25 January 1949) is an English geneticist, former President of the Royal Society and Chief Executive and Director of the Francis Crick Institute. He was awarded the 2001 Nobel Prize in Physiology or Medicine along with Leland Hartwell and Tim Hunt for their discoveries of protein molecules that control the division of cells in the cell cycle. Early life and education Nurse's mother went from London to Norwich, Norfolk and lived with relatives while awaiting Paul's birth (at the age of 18) in order to hide illegitimacy. For the rest of their lives, his maternal grandmother pretended to be his mother, and his mother pretended to be his sister. Paul was brought up by his grandparents (whom he took to be his parents) in North West London. He was educated at Lyon Park school in Alperton and Harrow County Grammar School. He received his BSc degree in biology in 1970 from the University of Birmingham and his PhD degree in 1973 from the University of East Anglia for research on Candida utilis. He then pursued postdoctoral work at the University of Bern, the University of Edinburgh and the University of Sussex. Nurse did not know that his "sister" was in fact his mother until he was in his 50s: his application for a green card for US residency while president of Rockefeller University was, to his surprise, rejected, despite his being a Nobel Prize winner, president of a university and a knight; this was because he had submitted a short-form UK birth certificate which did not name his parents. When he applied for a full birth certificate he discovered the truth, to his astonishment. Career and research Nurse continued his postdoctoral research at the laboratory of Murdoch Mitchison at the University of Edinburgh for the next six years (1973–1979). Beginning in 1976, Nurse identified the gene cdc2 in fission yeast (Schizosaccharomyces pombe). This gene controls the progression of the cell cycle from G1 phase to S phase and the transition from G2 phase to mitosis. In 1987, Nurse identified the homologous gene in human, Cdk1, which codes for a cyclin dependent kinase. Working in fission yeast, Nurse identified the gene cdc2, which controls the transition from G1 to S, when the cell grows in preparation for the duplication of DNA, and G2 to M, when the cell divides. With his postdoc Melanie Lee, Nurse also