been referred to this species. At about long, H. rossicus was the largest species of Hesperornis and among the largest hesperornithines, slightly smaller than the large Canadian genus Canadaga. Aside from its large size and different geographic location, H. rossicus differs from other Hesperornis in several features of the lower leg and foot, including a highly flattened metatarsus. In 2002, Martin and Lim formally recognized several new species for remains that had previously been unstudied or referred without consideration to previously named North American hesperornithines. These include the very small H. mengeli and H. macdonaldi, the slightly larger H. bairdi, and the very large H. chowi, all from the Sharon Springs member of the Pierre Shale Formation in South Dakota and Alberta, 80.5 million years ago. In addition, there are some unassigned remains, such as SGU 3442 Ve02 and LO 9067t and bones of an undetermined species from Tzimlyanskoe Reservoir near Rostov. The former two bones are probably H. rossicus; some remains assigned to that species in turn seem to belong to the latter undetermined taxon. Paleobiology Hesperornis was primarily marine, and lived in the waters of such contemporary shallow shelf seas as the Western Interior Seaway, the Turgai Strait, and the North Sea, which then were subtropical to tropical waters, much warmer than today. However, some of the youngest known specimens of Hesperornis have been found in inland freshwater deposits of the Foremost Formation, suggesting that some species of Hesperornis may have eventually moved, at least partially, away from a primarily marine habitat. Additionally, the species H. altus comes from the freshwater deposits at the base of the Judith River Formation. Traditionally, Hesperornis is depicted with a mode of locomotion similar to modern loons or grebes, and study of their limb proportions and hip structure has borne out this comparison. In terms of limb length, shape of the hip bones, and position of the hip socket, Hesperornis is particularly similar to the common loon (Gavia immer), probably exhibiting a very similar manner of locomotion on land and in water. Like loons, Hesperornis were probably excellent foot-propelled divers, but might have been ungainly on land. Like loons, the legs were probably encased inside the body wall up to the ankle, causing the feet to jut out to the sides near the tail. This would have prevented them from bringing the legs underneath the body to stand, or under the center