(Ellenberger, 1970) G. moshoeshoei (Ellenberger, 1970) G. olonensis Lapparent & Monetnat, 1967 G. palissyi (Gand, 1976) G. paulstris (Ellenberger, 1970) G. perriauxi (Demathieu & Gand, 1972) G. princeps (Ellenberger, 1970) G. protocrassidigitus (Ellenberger, 1970) G. rapidus (Ellenberger, 1974) G. romanovskyi (Gabunia & Kurbatov) G. quthingensis (Ellenberger, 1974) G. rectilineus (Ellenberger, 1970) G. sabinensis (Gand & Pellier, 1976) G. sassendorfensis (Kuhn, 1958) G. sauclierensis Demathieu & Sciau, 1992 G. schlauersbachensis (Weiss, 1934) G. socialis (Ellenberger, 1970) G. ssatoi Yabe, Inai, & Shikama, 1940 G. tenuis Hitchcock, 1858 G. toscanus (Huene, 1941) G. variabilis Lapparent & Monetnat, 1967 Paleopathology Fossil tracks can be informative about theropod pathologies but apparently pathological traits may be due to unusual behaviors. Sandstone stratum dating to the Norian in southern Wales preserves tracks of an individual with a deformed digit III attributed to the ichnogenus Anchisauripus. The distal end of the digit was consistently flexed. However, this apparent pathology could be caused by the animal rotating the tip of that digit when lifting the foot. Occurrences Grallator-type footprints have been found in formations dating from the Early Triassic through to the early Cretaceous periods. They are found in the United States, Canada, Europe, Australia, Brazil (Sousa and Santa Maria Formations) and China, but are most abundant on the east coast of North America, especially the Triassic and Early Jurassic formations of the northern part of the Newark Supergroup. Newark Supergroup tracks The most famous, and archetypal tracks that conform to the Grallator type are those found on the East Coast of North America, specifically from the Late Triassic to Early Jurassic Newark Supergroup. These footprints were likely made by an unidentified, primitive dinosaur similar to Coelophysis. The Newark Supergroup footprints show digits II, III and IV, but no trace of the shorter digits I and V which would likely have been present in a dinosaur of this stage. The outer two digits would have been stubby and ineffective, not touching the ground during walking or running. Despite losing most of their effectiveness, dinosaur evolution had not yet removed these digits to fully streamline the foot. This is known because rare specimens are found with traces of these outer digits. Digits II, III and IV have 3, 4 and 5 phalanges respectively, giving Grallator a ?-3-4-5-? digital formula. Although the Newark Supergroup Grallator tracks were made by a bipedal saurischian dinosaur, they can easily be mistaken for those