Ayla in the purpose Mut has for her. Physical traits Jondalar is portrayed as masculine and virile by Auel. Relationships Dalanar Dalanar is leader of the First Cave of the Lanzadonii, a splinter colony located in western Switzerland and former spouse of Marthona, now the former leader of the Ninth Cave of the Zelandonii. Dalanar and Marthona had a complicated and deep love affair about which the singers and story tellers still speak of with reverence and perhaps a touch of envy. As revealed in The Shelters of Stone, Dalanar is so close in physical resemblance to Jondalar that Ayla found herself momentarily confused by and responding physically (sexually) to his nearness. He is still good friends with Marthona, and leads his people to camp near the Ninth Cave encampment at the Zelandonii summer meeting central to the fifth book. Like Jondalar, Dalanar is a master flint knapper, but like his ex-mate Marthona, has a strong leadership drive, and need to assert himself in such a capacity. That need to express his opinion and corollary differences of opinion with Marthona is the best and only description in his back story as to why he and Marthona split apart and he came to lead a colony group from the Ninth Cave off to the east while Jondalar was just a toddler. Why the splinter colony (from the ninth cave) he founded is named the Lanzadonii vs. Zelandonii is never adequately explained, though it is clear in several scenes of The Shelters of Stone that the Lanzadonii are still considered to belongpart of the greater peopleat least in the minds of several groups of leaders and Zeladonii shamans. Thonolan Thonolan, the younger brother of Jondalar, is handsome, dark haired and significantly shorter than his very tall brother, but ready with a laugh, frequent smiles and constantly joking. He is fifteen to Jondalar's eighteen when he decides to undertake "his journey" to the mouth of the Great Mother River, and Jondalar decides to tag along to protect his younger sibling, whom he loves dearly. The second book, The Valley of Horses introduces Jondalar and Thonolan traveling from the lands of the Lanzadonii, in today's western Switzerland, across the great glaciers capping the Alps into the upper Danube valley (Germany or Austria, given fictional license and Ice Age glacier sizes) accompanying his younger impetuous and head-strong brother Thonolan on his journey, putatively to reach