Carians to join the rebellion. Furthermore, seeing the spread of the rebellion, the kingdoms of Cyprus also revolted against Persian rule without any outside persuasion. For the next three years, the Persian army and navy were fully occupied with fighting the rebellions in Caria and Cyprus, and Ionia seems to have had an uneasy peace during these years. At the height of the Persian counter-offensive, Aristagoras, sensing the untenability of his position, decided to abandon his position as leader of Miletus, and of the revolt, and he left Miletus. Herodotus, who evidently has a rather negative view of him, suggests that Aristagoras simply lost his nerve and fled. By the sixth year of the revolt (494 BC), the Persian forces had regrouped. The available land forces were gathered into one army, and were accompanied by a fleet supplied by the re-subjugated Cypriots, and the Egyptians, Cilicians and Phoenicians. The Persians headed directly to Miletus, paying little attention to other strongholds, presumably intending to tackle the revolt at its centre. The Ionians sought to defend Miletus by sea, leaving the defense of Miletus to the Milesians. The Ionian fleet gathered at the island of Lade, off the coast of Miletus. The Persians were uncertain of victory at Lade, so attempted to persuade some of the Ionian contingents to defect. Although this was unsuccessful at first, when the Persians finally attacked the Ionians, the Samian contingent accepted the Persian offer. As the Persian and Ionian fleets met, the Samians sailed away from the battle, causing the collapse of the Ionian battle line. Although the Chian contingent and a few other ships remained, and fought bravely against the Persians, the battle was lost. With defeat at Lade, the Ionian Revolt was all but ended. The next year, the Persians reduced the last rebel strongholds, and began the process of bringing peace to the region. The Ionian Revolt constituted the first major conflict between Greece and the Persian Empire, and as such represents the first phase of the Greco-Persian Wars. Although Asia Minor had been brought back into the Persian fold, Darius vowed to punish Athens and Eretria for their support for the revolt. Moreover, seeing that the myriad city states of Greece posed a continued threat to the stability of his empire, he decided to conquer the whole of Greece. In 492 BC, the first Persian invasion of Greece, the next phase of