they affect behaviour. To put this another way, natural selection does not directly select for true beliefs, but rather for advantageous behaviours. Plantinga distinguished the various theories of mind-body interaction into four jointly exhaustive categories: epiphenomenalism, where behaviour is not caused by beliefs. "if this way of thinking is right, beliefs would be invisible to evolution" so P(R|N&E) would be low or inscrutable Semantic epiphenomenalism, where beliefs have a causative link to behaviour but not by virtue of their semantic content. Under this theory, a belief would be some form of long-term neuronal event. However, on this view P(R|N&E) would be low because the semantic content of beliefs would be invisible to natural selection, and it is semantic content that determines truth-value. Beliefs are causally efficacious with respect to behaviour, but maladaptive, in which case P(R|N&E) would be low, as R would be selected against. Beliefs are causally efficacious with respect to behaviour and also adaptive, but they may still be false. Since behaviour is caused by both belief and desire, and desire can lead to false belief, natural selection would have no reason for selecting true but non-adaptive beliefs over false but adaptive beliefs. Thus P(R|N&E) in this case would also be low. Plantinga pointed out that innumerable belief-desire pairs could account for a given behaviour; for example, that of a prehistoric hominid fleeing a tiger: Perhaps Paul very much likes the idea of being eaten, but when he sees a tiger, always runs off looking for a better prospect, because he thinks it unlikely the tiger he sees will eat him. This will get his body parts in the right place so far as survival is concerned, without involving much by way of true belief. ... Or perhaps he thinks the tiger is a large, friendly, cuddly pussycat and wants to pet it; but he also believes that the best way to pet it is to run away from it. ... Clearly there are any number of belief-cum-desire systems that equally fit a given bit of behaviour. Thus, Plantinga argued, the probability that our minds are reliable under a conjunction of philosophical naturalism and naturalistic evolution is low or inscrutable. Therefore, to assert that naturalistic evolution is true also asserts that one has a low or unknown probability of being right. This, Plantinga argued, epistemically defeats the belief that naturalistic evolution is true and that ascribing truth to