a somewhat larger and likewise unnamed azhdarchid, with a wingspan of ; and apparently small pteranodontids have been found as well. The robust, flightless, and possibly herbivorous avialan or dromaeosaurid Balaur, which had two enlarged claws on each foot, represents another highly specialized component of the fauna. The ecosystem contained a number of insular dwarfs, namely the titanosaurs Magyarosaurus and Paludititan, the hadrosaurid Telmatosaurus, and the iguanodontian Zalmoxes. Along with the nodosaurid Struthiosaurus, various small, fragmentary maniraptorans were present: Bradycneme, Elopteryx, and Heptasteornis. Crocodilian remains, belonging to the genera Allodaposuchus, Doratodon, and Acynodon have also been found. Non-archosaurian components include the kogaionid multituberculate mammals Kogaionon, Barbatodon, Litovoi tholocephalos, and Hainina; lizards such as the teiid Bicuspidon and the paramacellodid Becklesius; an unnamed madtsoiid snake; and the lissamphibians Albanerpeton, Eodiscoglossus, and Paradiscoglossus''. During the Maastrichtian, southern Europe was an archipelago. The members of the Hațeg Island ecosystem lived on a landmass known as the Tisia–Dacia Block, of which the Hațeg Basin was a small part. This landmass was about in area, and was separated from other terrestrial terrains by stretches of deep ocean in all directions by . Being located at 27°N, the island was located farther south than the present-day latitude of 45°N; the climate was likely subtropical, with distinct dry and wet seasons, and an average temperature of about . The environment consisted of various alluvial plains, wetlands, and rivers, surrounded by woodlands dominated by ferns and angiosperms. Paleosols indicate a relatively dry Cretaceous climate, with an annual precipitation of less than . See also List of pterosaur genera Timeline of pterosaur research Pterosaur size References External links Azhdarchids Late Cretaceous pterosaurs of Europe Geography of Transylvania Maastrichtian life Fossil taxa described in 2002 Hațeg fauna Apex predators