relations between Scythian and Bosporan royalty. In the 4th century BC, the Bosporan kingdom became the main supplier of grains to Greece partly because of the Peloponnesian War which was raging in the latter region, which intensified the grains trade between the Scythians and the Greeks, with the Scythians becoming the principal middlemen in the supply of grains to the Bosporan kingdom: while most of the grains that the Scythians sold to the Greeks was produced by the agricultural populations in the northern forest steppe, the Scythians themselves were also trying to produce more grains within Scythia itself, which was a driving force behind the sedentarisation of many of the hitherto nomadic Scythians; the process of Scythian sedentarisation thus was most intense in the regions adjacent to the Bosporan cities in eastern Crimea. The Scythian royalty and aristocracy obtained enormous profits from this grains trade, and this period saw Scythian culture not only thriving, with most known Scythian monuments dating from then, but also rapidly undergoing significant Hellenisation. The city of the Kamianka site remained the political, industrial and commercial capital of Scythian during the 4th and early 3rd centuries BC, during which time the Scythians founded a new settlement at which functioned as the main administrative, commercial and industrial centre of the lower Don river and northern Lake Maeotis areas and was also the residence of local Scythian lords. The main burial centre of the Scythians during this period was located in the Nikopol and Zaporizhzhia region on the lower Dnipro, where were located the Solokha, , Krasnokutsk and kurgans. Rich burials, such as, for example, the , attest of the wealth acquired from the grains trade by the Scythian aristocracy of the 4th century BC, who were progressively buried with more, relatives, retainers, and grave goods such as gold and silver objects, including Greek-manufactured toreutics and jewellery; the Scythian commoners however did not obtain any revenue from this trade, and luxury items are absent from their burials. Despite the pressure of some smaller and isolated Sarmatian groups in the east, the period remained largely and unusually peaceful and the Scythian hegemony in the Pontic steppe remained undisturbed, with the Scythian nomads continuing to form the bulk of the northern Pontic region's population. The most famous Scythian king of the 4th century BC was Ateas, who was the successor and possibly the son of the Scythian king buried at