sex ratio is the ratio of the allotted investment between the sexes, and is sometimes referred to as Fisherian sex ratios. However, there are many examples of organisms that do not demonstrate the expected 1:1 ratio or the equivalent investment ratio. The idea of equal allocation fails to explain these expected ratios because it assume that relatives do not interact with one another, and that the environment has no effect. Interactions between relatives W.D. Hamilton hypothesized that non-Fisherian sex ratios can result when relatives interact with one another. He argued that if relatives experienced competition for resources, or benefited from the presence of other relatives, then sex ratios would become skewed. This led to a great deal of research on whether competition or cooperation between relatives results in differential sex ratios that do not support Fisher's principle. Local resource competition Local resource competition (LRC) was first hypothesized by Anne Clark. She argued that the African bushbaby (Otolemur crassicaudatus) demonstrated a male-biased sex ratio because daughters associated with mothers for longer periods of time than did sons. Since sons disperse further from the maternal territory than do daughters, they do not remain on the territories and do not act as competitors with mothers for resources. Clark predicted that the effect of the LRC on sex allocation resulted in a mother investing preferentially in male offspring to reduce competition between daughters and herself. By producing more male offspring that disperse and do not compete with her, the mother will have a greater fitness than she would if she had produced the ratio predicted by the equal investment theory. Further research has found that LRC may influence the sex ratio in birds. Passerine birds demonstrate largely daughter-based dispersal, while ducks and geese demonstrate mainly male-based dispersal. Local resource competition has been hypothesized to be the reason that passerine birds are more likely to be female, while ducks and geese are more likely to have male offspring. Other studies have hypothesized that LRC is likely to influence sex ratios in roe deer, as well as primates. Consistent with these hypotheses, the sex-ratios in roe deer and several primates have been found to be skewed towards the sex that does not compete with mothers. Local mate competition Local mate competition (LMC) can be considered a special type of LRC. Fig wasps lay fertilized eggs within figs, and no females disperse. In some species, males are