leaves, which may also be seen after bushfire, are broader and serrated. Leaf undersides have fine white hairs in the case of the varieties spinulosa and collina and pale brown in cunninghamii and neoanglica. The distinctive inflorescences or flower spikes occur over a short period through autumn and early winter. A spike may contain hundreds or thousands of individual flowers, each of which consists of a tubular perianth made up of four united tepals, and one long wiry style. Characteristic of the taxonomic section in which it is placed, the styles are hooked rather than straight. The style ends are initially trapped inside the upper perianth parts, but break free at anthesis. In Banksia spinulosa the spikes are cylindrical, about wide and tall, yellow to golden orange in colour, with styles varying from yellow to pink, maroon, or black. Styles of various colours may be found within metres of each other in some areas such as in the Georges River National Park, and Catherine Hill Bay, while other populations may have uniformly black, red or gold styles. Though not terminal, the flower spikes are fairly prominently displayed. Partly emerging from the foliage, they arise from two- to three-year-old stem nodes. The hairpin banksia's infructescence is a typical Banksia cone-like structure, with up to 100 crowded embedded follicles which are 1–2.4 centimetres (⅓–1 in) in diameter; these generally remain closed until burnt by bushfire. The nonlignotuberous subspecies cunninghamii is killed by fire and regenerates from seed, while the others regenerate from buds around the base of the lignotuber. Old flower spikes fade to brown, then grey with age. Old flower parts usually persist for a long time, giving the infructescence a hairy appearance. In Central and North Queensland, old cones of both var. spinulosa and var. collina are generally bare. Taxonomy The first known specimens of B. spinulosa were collected near Sydney by John White, Surgeon General to the British colony of New South Wales, sometime between 1788 and 1793. He called it "prickly-leaved banksia", though this name has fallen out of use. It is uncertain exactly when he first collected the species; it may have been before 1790, as there is speculation that a sketch in his 1790 Journal of a Voyage to New South Wales is of a B. spinulosa infructescence. Text accompanying the figure states English botanist James Edward Smith later tentatively attributed this figure to B. spinulosa: