the Mōri family for the duration of the Edo period. Because the shogunate frequently confiscated domains whose daimyo were unable to produce heirs, the Mōri daimyo created four subordinate han ruled by branches of the family: Iwakuni han: 60,000 koku, ruled by descendants of Kikkawa Hiroie. Chōfū han: 50,000 koku, ruled by descendants of Mōri Hidemoto. Tokuyama han: 40,000 koku, ruled by descendants of Mōri Naritaka. Kiyosue han: 10,000 koku, ruled by descendants of Mōri Mototomo. During the Edo period, the main branch died out in 1707, after which heirs were adopted from the Chōfu branch, which also became extinct in 1751. The family then continued through the Kiyosue branch. The Mōri daimyo, as with many of his counterparts throughout Japan, was assisted in the government of his domain by a group of karō, or domain elders. There were two kinds of karō in Chōshū: hereditary karō (whose families retained the rank in perpetuity) and the "lifetime karō", whose rank was granted to an individual but could not be inherited by his son. The hereditary karō were either members of minor branches of the Mōri family, or members of related families such as the Shishido and the Fukuhara, or descendants of Mōri Motonari's most trusted generals and advisors such as the Mazuda, the Kuchiba and the Kunishi. The lifetime karō were middle or lower samurai who displayed great talent in economics or politics and were promoted to karō by the daimyō. One such person was the great reformer Murata Seifu. List of daimyōs Mōri clan (Tozama, 369,000 koku), 1600–1871 Simplified family tree of the main Mōri line (Lords of Chōshū) Mōri Motonari (1497–1571) Takamoto (1523–1563) I. Terumoto, 1st Lord of Chōshū (cr. 1600) (1553–1625; r. 1600–1623) II. Hidenari, 2nd Lord of Chōshū (1595–1651; r. 1623–1651) III. Tsunahiro, 3rd Lord of Chōshū (1639–1689; r. 1651–1682) IV. Yoshinari, 4th Lord of Chōshū (1668–1694; r. 1682–1694). V. Yoshihiro, 5th Lord of Chōshū (1673–1707; r. 1694–1707) Naritaka, 1st Lord of Tokuyama (1602–1679) Mototsugu, 3rd Lord of Tokuyama (1667–1719) Hirotoyo, 5th Lord of Tokuyama (1705–1773) Nariyoshi, 7th Lord of Tokuyama (1750–1828) Hiroshige, 8th Lord of Tokuyama (1777–1866) XV. Motonori, 15th Lord of Chōshū, 1st Prince (1839–1896; r. 1869, Governor of Hagi 1869–1871, family head 1871–1896, created 1st Prince 1884) Motoaki, 29th family head, 2nd Prince (1865–1938; 29th family head and 2nd Prince 1896–1938) Motomichi, 30th family head, 3rd Prince (1903–1976; 30th family head 1938–1976, 3rd