for many years. During this period his interest in the role of insects in transmission of disease developed, which would continue to be a major topic of research. In May 1899 Nuttall travelled to Cambridge at the invitation of Clifford Allbutt, Regius Professor of Physic at the University, and gave a series of lectures in bacteriology. In 1900 he was appointed university lecturer in bacteriology and preventive medicine, and would be based in Cambridge for the rest of his life. He founded and edited the Journal of Hygiene, the first volume being published in 1901. His research at this period was in two main areas, studies of blood, in particular immune reactions, and studies on transmission of disease by arthropods, in particular mosquitoes and malaria with Arthur Shipley. In 1904 Nuttall and Shipley were both elected Fellows of the Royal Society. In the same year he and Patrick Manson established in Cambridge the first Diploma in Tropical Medicine and Hygiene, which continued until 1933. In 1906 he was elected the first Quick Professor of Biology at Cambridge, a chair established for the study of protozoology. He built a substantial team within the Quick Laboratory working on many areas of parasitology. A major topic was piroplasmosis and related malaria-like parasites transmitted by ticks, mainly in dogs, but also in other animals including humans. The number of published papers on parasitology was steadily increasing, and in 1908 Nuttall founded the journal Parasitology, initially as a supplement to the Journal of Hygiene, but soon as a separate journal. Cox (2009) argues that the founding of the journal was a key event in the development of parasitology as a discipline in its own right. In the same year Nuttall co-edited an important book on the bacteriology of diphtheria with Graham-Smith. Also in 1908 he was elected a felllow of Magdalen College. During World War I he began to investigate lice. This began in response to the practical problems with lice in the troops, but developed over the next few years into research on their biology and role in disease. In 1919 Nuttall appealed for funds to create an Institute for Parasitical Research in Cambridge. Percy Molteno and his wife donated the sum of £30,000 for this purpose. The Molteno Institute for Research in Parasitology was opened in 1921 with Nuttall as its first director. The Quick Department was transferred to the Molteno Institute. His