spores. Mycangia in leaf-rolling weevils Females of the leaf-rolling weevil in the genus Euops (Coleoptera: Attelabidae) store symbiotic fungi in the mycangia, which is between the first ventral segment of the abdomen and the thorax. Different from ovipositor-associate mycangia in woodwasps, lizard beetles, and ship-timber beetles, mycangia of leaf-rolling weevils is a pair of spore incubators at the anterior end of the abdomen. This mycangium is formed by the coxa and the metendosternite at the posterior end of the thorax. Mycangia in stag beetles Mycangia of the stag beetles (Coleoptera: Lucanidae) were discovered in Japan only this century. This ovipositor-associated mycangium is located in a dorsal fold of the integument between the last two tergal plates of the adult females. It has been examined in many species. A female everts the mycangium for the first time soon after eclosion; this is to retrieve the symbionts left by the larva on the pupal chamber when it emptied its gut before pupating. Later, when ovipositing, she everts it to pass on the inoculum to the next generation. References Symbiosis Insect anatomy