Anne Casimir Pyramus (or Pyrame) de Candolle (20 February 1836, Geneva – 3 October 1918, Chêne-Bougeries) was a Swiss botanist, the son of Alphonse Pyramus de Candolle. Early life and education He studied chemistry, physics and mathematics in Paris (1853–57), later spending time in England, where he met with Miles Berkeley. In 1859 he visited Algeria, and during the following year, continued his education in Berlin. Afterwards, he returned to Geneva as an assistant and colleague to his father. He married Anna-Mathilde Marcet and they had four children: Raymond Charles de Candolle (1864–1935), Florence Pauline Lucienne de Candolle (1865–1943), Richard Émile Augustin de Candolle (1868–1920) and Reyne Marguerite de Candolle (1876–1958). Career In the field of plant systematics, he used criteria such as stem structure and/or leaf arrangement as a basis of anatomical criteria. As a plant physiologist, he conducted investigations on the movement of leaves, the curling of tendrils, the effect of low temperatures on seed germination and the influence of ultraviolet radiation on flower formation. He was particularly interested in the botanical family Piperaceae. He continued work on , a project begun by his father, and was co-editor of the (Geneva). He held honorary degrees (doctor honoris causa) from the universities of Rostock, Geneva, Aberdeen and Uppsala. References External links 20th-century Swiss botanists Plant physiologists 1836 births 1918 deaths Scientists from Geneva 19th-century Swiss botanists