more restricted; Thalassodromeus may have been better at handling relatively large, struggling prey than its relative, Tupuxuara, which had a more lightly built skull. Witton stressed that more studies of functional morphology would have to be done to illuminate the subject and speculated that Thalassodromeus might have been a raptorial predator, using its jaws to subdue prey with strong bites; its concave palate could help it swallow large prey. Pêgas and Kellner presented a reconstruction of the mandibular muscles of T. sethi at a conference in 2015. They found that its well-developed jaw muscles differed from those of the possible dip-feeder Anhanguera and the terrestrially stalking azhdarchids, indicating that T. sethi had a strong bite force. In 2018, Pêgas and colleagues agreed that Thalassodromeus blade-like, robust jaws indicated that it could have used them to strike and kill prey, but they thought that biomechanical work was needed to substantiate the idea. They found (unlike Witton) that Thalassodromeus had a reinforced jaw joint and robust jaw muscles, but more work was needed to determine its dietary habits. According to Pêgas and colleagues, the articulation between T. sethi articular and quadrate bones (where the lower jaw connected with the skull) indicates a maximum gape of 50degreessimilar to the 52-degree gape inferred for Quetzalcoatlus. Locomotion In a 2002 comment on the original description of T. sethi, engineer John Michael Williams noted that although Kellner and Campos had mentioned that the large crest might have interfered aerodynamically during flight, they had not elaborated on this point and had compared the pterosaur with a bird one-fifth its size. He suggested that Thalassodromeus used its crest to balance its jaws, with the head changing attitude depending on the mode of locomotion. Williams speculated that the crest would be inflatable with blood and presented varying air resistance, which he compared to a handheld fan; this would have helped the animal change the attitude of the head during flight (and during contact with water), keeping it from rotating without powerful neck muscles. The crest would have made long flights possible, rather than interfering; Williams compared it with the spermaceti in the head of the sperm whale, stating it is supposedly used to change buoyancy through temperature adjustment. Kellner and Campos rejected the idea of an inflatable crest, since its compressed bones would not allow this; they did not find the sperm-whale analogy convincing in relation to flying animals,