The most widespread plants were Cycadales and the conifer Brachyphyllum. Other pterosaurs from the Romualdo Formation include Anhanguera, Araripedactylus, Araripesaurus, Brasileodactylus, Cearadactylus, Coloborhynchus, Santanadactylus, Tapejara, Tupuxuara, Barbosania, Maaradactylus, Tropeognathus, and Unwindia. Thalassodromines are known only from this formation, and though well-preserved postcranial remains from there have been assigned to the group, they cannot be assigned to genus due to their lack of skulls. Dinosaur fauna includes theropods like Irritator, Santanaraptor, Mirischia, and an indeterminate unenlagiine dromaeosaur. The crocodyliforms Araripesuchus and Caririsuchus, as well as the turtles Brasilemys, Cearachelys, Araripemys, Euraxemys, and Santanachelys, are known from the deposits. There were also clam shrimps, sea urchins, ostracods, and molluscs. Well-preserved fish fossils record the presence of hybodont sharks, guitarfish, gars, amiids, ophiopsids, oshuniids, pycnodontids, aspidorhynchids, cladocyclids, bonefishes, chanids, mawsoniids and some uncertain forms. Pêgas and colleagues noted that pterosaur taxa from the Romualdo Formation had several species: two of Thalassodromeus, two of Tupuxuara, and up to six species of Anhanguera. It is possible that not all species in each taxon coexisted in time (as has been proposed for the pteranodontids of the Niobrara Formation), but there is not enough stratigraphic data for the Romualdo Formation to test this. See also List of pterosaur genera Timeline of pterosaur research References Bibliography Tapejaromorphs Early Cretaceous pterosaurs of South America Albian life Cretaceous Brazil Fossils of Brazil Romualdo Formation Fossil taxa described in 2002 Taxa named by Alexander Kellner