The Satyrini is one of the tribes of the subfamily Satyrinae. It includes about 2200 species and is therefore the largest tribe in the subfamily which comprises 2500 species. Distribution Satyrini butterflies have a worldwide distribution, but the distribution pattern differs between subtribes. Some subtribes are almost restricted to a single biogeographic region, such as the Pronophilina, which is found only in Andean cloud forests from Venezuela to Bolivia. Biology The larval food plants of many species in this tribe are grasses, i.e. Poaceae. It is considered that the Satyrini diversified at about the same time as the grasses did, and that the radiation of the tribe is therefore closely related to the evolution of the grasses. In contrast, the tribe has a few genera which show uncommon feeding preferences. Three genera, Euptychia, Ragadia and Acrophtalmia, feed on Lycopsida, and moreover, some species of Euptychia have been reported to feed on mosses of Neckeraceae. This is interesting cases as these genera are not considered to be very closely related to each other. Taxonomy The taxonomy of the subfamily Satyrinae has recently undergone a significant revision. showed that most of the traditionally well known group are paraphyletic or polyphyletic and developed a new taxonomy, based on molecular phylogenetic. The classification scheme transferred many genera traditionally placed in the Elymniini to the Satyrini, and later studies have largely followed this classification. Thus, the Satyrini is now considered to include the genera previously placed in the Elymniini, such as Lethe, Pararge and Mycalesis. However, the phylogenetic statuses of some subtribes within the tribe are still unclear and the revision of the classification is likely to continue. According to , the Satyrini includes the following subtribes and genera: Parargina Moore, 1893 Chonala Moore, 1893 Kirinia Westwood, 1841 Lasiommata Tutt, 1896 Lopinga Moore, 1893 Orinoma Gray, 1846 Pararge Hübner, 1819 Rhaphicera Butler, 1867 Tatinga Moore, 1893 Nosea Moore, 1893 Lethina Reuter, 1896 Aphysoneura Karsch, 1894 Lethe Hübner, 1819 Neope Moore, 1866 Ninguta Moore, 1892 Ptychandra Felder & Felder, 1861 Mandarinia Leech, 1892 Mycalesina Reuter, 1896 Lohora Moore, 1880 Brakefieldia Aduse-Poku, Lees & Wahlberg, 2016 Telinga Moore, 1880 Heteropsis Westwood, 1850 Culapa Moore, 1879 Mydosama Moore, 1880 Mycalesis Hübner, 1818 Devyatkinia Monastyrskii & Uémura, 2016 Hallelesis Condamin, 1961 Bicyclus Kirby, 1871 Eritina Miller, 1968 Coelites Westwood, 1850 Erites Westwood, 1851 Orsotriaena Wallengren, 1858 Zipaetis Hewitson, 1863 Ragadiina Herrich-Schäffer, 1864 Acrophtalmia Felder & Felder, 1861 Acropolis Hemming,