Kellner and Campos in 1988 based on fossils from the same formation. In the view of Martill and Naish, the differences between these genera (including two species of Tupuxuara, T. longicristatus and T. leonardii) were due to ontogeny (changes during growth) and compression of the fossils; Thalassodromeus was simply an older, larger, and better-preserved individual. This idea was rejected by Kellner and Campos in 2007, who pointed out these species had differences in features other than their crests. They also noted that one specimen of Tupuxuara had a larger skull than Thalassodromeus (measured from the tip of the premaxilla to the back of the squamosal bone), despite Martill and Naish's contention that the latter was an older individual. Kellner and Campos' view has since been accepted by other researchers, including Martill and Naish. Veldmeijer and colleagues assigned the front part of a mandible collected from the same formation to T. sethi in 2005. They concluded that although the two specimens differed in several details, the differences were not significant enough to base a new species on the mandible, and that the new specimen filled in the gap of Kellner and Campos' T. sethi skull reconstruction. Palaeontologists Jaime A. Headden and Herbert B. N. Campos coined the new binomial Banguela oberlii, based on their reinterpretation of the jaw tip as belonging to a toothless member of the family Dsungaripteridae, in 2014. The generic name is Portuguese for "toothless" and the specific name honours private collector Urs Oberli, who had donated the specimen to the Naturmuseum St. Gallen (where it is catalogued as NMSG SAO 25109). Headden and Campos interpreted the tip of T. sethi lower jaw as downturned; this and other features distinguished it from Banguela. In their 2018 re-description of the further-prepared T. sethi holotype skull, palaeontologists Rodrigo V. Pêgas, Fabiana R. Costa, and Kellner assigned B. oberlii back to Thalassodromeus while recognising it as a distinct species, and thereby created the new combination T. oberlii. Pêgas and colleagues also rejected the theory that the lower jaw of T. sethi was downturned, and reinterpreted the frontmost piece of the lower jaw to have connected directly with the subsequent piece (with no gap). In 2015 palaeontologists Gerald Grellet Tinner and Vlad A. Codrea named a new species, T. sebesensis, based on what they interpreted as part of a cranial crest in a concretion found near the Sebeș River in Romania. The