the bombing, the French Directorate-General for External Security was informed by their British counterpart that the UK suspected the Libyans to be behind the bombing. According to a CIA analysis dated 22 December 1988, several groups were quick to claim responsibility in telephone calls in the United States and Europe: A male caller claimed that a group called the "Guardians of the Islamic Revolution" had destroyed the plane in retaliation for Iran Air Flight 655 being shot down by US forces in the Persian Gulf the previous July. A caller claiming to represent the Islamic Jihad Organization told ABC News in New York that the group had planted the bomb to commemorate Christmas. Another caller claimed the plane had been downed by Mossad, the Israeli intelligence service. The list's author noted, "We consider the claims from the Guardians of the Islamic Revolution as the most credible one received so far," but the analysis concluded, "We cannot assign responsibility for this tragedy to any terrorist group at this time. We anticipate that, as often happens, many groups will seek to claim credit." Muammar Gaddafi took responsibility for the Lockerbie bombing and paid compensation to the victims' families in 2003, though he maintained that he had not ordered the attack. On 22 February 2011, during the Libyan Civil War, former Minister of Justice Mustafa Abdul Jalil stated in an interview with the Swedish newspaper Expressen that Gaddafi had personally ordered the bombing. Jalil claimed to possess "documents that prove [his allegations] and [that he is] ready to hand them over to the international criminal court." In December 2013, the original prime suspects in the bombing were revealed to have been the Popular Front for the Liberation of Palestine – General Command (PFLP-GC), a Syria-based group led by Ahmed Jibril. A flood of warnings immediately preceding the disaster had included one that read: 'team of Palestinians not associated with PLO intends to attack US targets in Europe. Time frame is present. Targets specified are Pan Am Airlines and US military bases.' Five weeks before this warning, Jibril's right-hand man, Haffez Dalkamoni, had been arrested in Frankfurt with a known bomb-maker, Marwen Khreesat. "Later US intelligence officials confirmed that members of the group had been monitoring Pan Am's facilities at Frankfurt airport. On Dalkamoni's account bombs made by Khreesat were at large somewhere." A deep-cover CIA agent was told by up to 15 high-level