authors said that this would extend the range in time and space for the genus Thalassodromeus considerably, creating a 42-million-year gap between the older South American species and the younger European species. Palaeontologist Gareth J. Dyke and a large team of colleagues immediately rejected the pterosaurian identification of the T. sebesensis fossil, instead arguing that it was a misidentified part of a plastron (lower shell) of the prehistoric turtle Kallokibotion bajazidi (named in 1923). The idea that the fragment belonged to a turtle had been considered and rejected by Grellet-Tinnera and Codrea in their original description. Grellet-Tinnera and Codrea denied the turtle identity suggested by Dyke and colleagues, noting that those researchers had not directly examined the fossil. Description The holotype (and only known skull) of Thalassodromeus sethi is one of the largest pterosaur skulls ever discovered. The entire skull is estimated to have been long; the bones were fused together, indicating adulthood. Based on related pterosaurs, its wingspan was , making Thalassodromeus the largest known member of its clade, Thalassodromidae. Of similar proportions, its skull was more heavily built than that of its relative Tupuxuara. Although the postcranial skeleton of Thalassodromeus is unknown, relatives had unusually short and blocky neck vertebrae, with well-developed front and hind-limbs that were almost equal in length (excluding the long wing-finger). The hindlimbs were eighty percent that of the forelimb length, a unique ratio among pterodactyloids (short-tailed pterosaurs). As a pterosaur, Thalassodromeus was covered with hair-like pycnofibres and had extensive wing membranes (which were extended by the wing finger). The skull of T. sethi had a streamlined profile, especially from the tip of the snout to the front edge of the nasoantorbital fenestra (opening which combined the antorbital fenestra in front of the eye with the bony nostril). The most conspicuous feature of the skull was the large crest, which ran along the upper edge from the tip of the snout and beyond the occiput at the back of the skull, almost doubling the length and height of the skull. With the exception of the pterosaur Tupandactylus imperator (whose crest consisted mainly of soft tissue), T. sethi had the proportionally largest cranial crest of any known vertebrate (75 percent of the skull's side surface). The crest was mainly formed by the premaxillae (the frontmost snout bones), frontal bones, parietal bones, and part of the supraoccipital bone. The premaxillae formed most of the crest, extending